EUCYCLOPINAE Kiefer, 1927


The subfamily Eucyclopinae presently includes 10/11 genera and about 185 species and subspecies, including the genus Austriocyclops, formerly included in the subfamily Cyclopinae, and recently moved to Eucyclopinae by Pospisil & Stoch (1977).



ISOCYCLOPS Alekseev, 2019

DEFAYEICYCLOPS Alekseev et Vaillant, 2013

MACROCYCLOPS Claus, 1893

EUCYCLOPS Claus, 1893

PARACYCLOPS Claus, 1893

ECTOCYCLOPS Brady, 1904

STYGOCYCLOPS Alekseev, 2019

TROPOCYCLOPS Kefer, 1927

AFROCYCLOPS Sars, 1927

THAUMASIOCYCLOPS Kiefer, 1930

OCHRIDACYCLOPS Kiefer, 1937

HOMOCYCLOPS (Herrick, 1882) *


* synonym of Macrocyclops (?)





MACROCYCLOPS Claus, 1893 (syn. Pachycyclops Sars G.O., 1914)


  • Macrocyclops albidus albidus (Jurine, 1820) [cosmopolitan;fresh waters, ground waters] [PDF]
  • Macrocyclops albidus oligolasius Kiefer, 1928 [cosmopolitan;fresh waters]
  • Macrocyclops albidus f. platysetus Fryer, 1957 [Africa; lakes]
  • Macrocyclops albidus albidus (Jurine, 1820)
  • Macrocyclops albidus oligolasius Kiefer, 1934
  • Macrocyclops albidus principalis Herbst, 1963 [Venezuela, Brazil, Colombia ; fresh waters, ephemeral wetland]
  • Macrocyclops ater (Herrick, 1882) [USA, Cuba; fresh waters] (syn. Homocyclops ater (Herrick, 1882) [PDF] *
  • Macrocyclops baicalensis Mazepova, 1961 [ Baikal; lakes]
  • Macrocyclops bistriatus (Koch, 1841)
  • Macrocyclops distinctus (Richard, 1887) [Europe, New Zealand, Indie, Korea; fresh waters]
  • Macrocyclops fuscus (Jurine, 1820) [cosmopolitan; fresh waters]
  • Macrocyclops monticola Ishida, 1994 [Japan; stream waters]
  • Macrocyclops neuter Kiefer, 1931 [Java, Sri Lanka, Birmania; fresh waters] (left) [PDF]

  • Macrocyclops oithonoides Roen, 1957
  • Macrocyclops oligolasius Kiefer, 1938
  • Macrocyclops signatus (Koch, 1841)
  • Macrocyclops tenuicornis (Claus, 1857) (syn. Macrocyclops signatus (Koch, 1841)


    * Suarez-Morales et al. (1996) included this species in the genus Homocyclops, by most authors considered as synonym of Macrocyclops. Reid (in litt.) confirmed that remarkable differences separate the nominate genera, which should be considered distinct taxa.



    PARACYCLOPS Claus, 1893 (syn. Platycyclops Sars G.O., 1914)


    The genus Paracyclops Claus, 1893 presently includes 33 species and subspecies, worldwide distributed, in different freshwater and brackishwater bodies, as well as occasionally in grounds waters.

    The taxonomy of the genus is still problematic because of the lack of complete early descriptions of widely distributed nominal species such as Paracyclops chiltoni (Thomson, 1882) and Paracyclops fimbriatus (Fischer, 1853) (Karaytug 1999; Karaytug and Boxshall 1998a; Dussart and Defaye 2006). Species of Paracyclops have been recorded from different types of freshwater habitats, including estuaries and semi-terrestrial habitats such as bromeliads (Karaytug and Boxshall 1998b); however, most are benthic forms (Karaytug 1999). The genus is distributed worldwide, including cold temperate latitudes and fully tropical areas, where the genus tends to be most speciose (from: Mercado-Salas & Suarez-Morales, 2009).


  • Paracyclops affinis (G.O. Sars, 1863) [cosmopolitan, except Australia; fresh waters]
  • Paracyclops abnobensis Kiefer, 1929
  • Paracyclops aioiensis Ito, 1957 [Japan; fresh waters] [PDF]
  • Paracyclops altissimus Karaytug, Defaye & Boxshall, 1998 [Africa; rivers] [PDF]
  • Paracyclops andinus Kiefer, 1957 [Peru, Argentina, lakes] [PDF] [PDF2]
  • Paracyclops baicalensis Mazepova, 1961 [Baikal lake]
  • Paracyclops bromeliacola Karaytug & Boxshall, 1998 [Brazil; bromeliads]
  • Paracyclops canadensis (Willey, 1934) [north America, Canada; lakes]

  • Paracyclops canthocarpoides (Fischer, 1851) (= Ectocyclops phaleratus (Koch, 1838)
  • Paracyclops carectum Reid, 1987 [Brazil; ponds, lakes, wet campos] [PDF]
  • Paracyclops chiltoni (Thomson, 1882) [Europe, USA, Russia, Brazil, Australia, New Zealand, Africa, Tahiti; surface, ground fresh waters] [PDF]
  • Paracyclops coperes Gurney, 1928 accepted as Ectocyclops coperes (Gurney, 1928)
  • Paracyclops crassicornis (Müller O.F., 1785) accepted as Paracyclops fimbriatus fimbriatus (Fischer, 1853)
  • Paracyclops dilatatus dilatatus Lindberg, 1952 [Caspian Sea]
  • Paracyclops dilatatus ivanegai Monchenko, 1977 [URSS: brackish waters]
  • Paracyclops eucyclopoides Kiefer, 1929 (syn. Australoeucyclops eucyclopoides (Kiefer, 1929)
  • Paracyclops fimbriatus fimbriatus (Fischer, 1853) [Europe, Asia; surface, ground fresh waters] (left)
  • Paracyclops fimbriatus andinus Lindberg, 1958
  • Paracyclops fimbriatus andinus Lindberg, 1958
  • Paracyclops fimbriatus bromeliarum Herbst, 1959
  • Paracyclops fimbriatus imminutus Kiefer, 1929
  • Paracyclops fimbriatus orientalis Alekseev, 1995
  • Paracyclops fimbriatus euchaetus Kiefer, 1939 [Africa, fresh water]
  • Paracyclops fimbriatus paropamisi Lindberg, 1960 [Afghanistan; pools]
  • Paracyclops finitimus Kiefer, 1929
  • Paracyclops fischeri Najam-un-Nisa, Mahoon & Irfan Khan, 1987
  • Paracyclops hardingi Karaytug & Boxshall, 1998 [Perù; Ecuador; fresh waters]
  • Paracyclops hirsutus Mercado-Salas & Suarez-Morales, 2009 [Mexico; fresh-waters] [PDF]
  • Paracyclops imminutus Kiefer, 1929 [Europe, Russia, Israel, Azores; fresh waters]
  • Paracyclops intermedius Tang & Knott, 2009 [Western Australia; ground waters] [PDF]
  • Paracyclops eucyclopoides Kiefer, 1929 (syn. Australoeucyclops eucyclopoides (Kiefer, 1929)
  • Paracyclops linderi (Lindberg, 1948) accepted as Australoeucyclops linderi (Lindberg, 1948
  • Paracyclops longispina Karaytug, Defaye & Boxshall, 1998 [Kenya; lakes] [PDF]
  • Paracyclops novenarius Reid, 1987 [Colombia; artificial containers]
  • Paracyclops oligarthrus (G.O. Sars, 1909) [Lake Tanganika]
  • Paracyclops orientalis Alekseev, 1995 [URSS; water reservoirs]
  • Paracyclops phaleratus (Koch, 1838) (syn. Platycyclops phaleratus (Koch, 1838)
  • Paracyclops pilosus Dussart, 1984 [Venezuela, Guyane; fresh waters] [PDF]
  • Paracyclops poppei (Rehberg, 1880) [Europe, Ukraine, central and south America, Africa ?, Asia, Israel; fresh waters]
  • Paracyclops punctatus Karaytug & Boxshall, 1998 [Brazil: bromeliads]
  • Paracyclops rehbergi Mahoon & Zia, 1985
  • Paracyclops reidae Karaytug & Boxshall, 1998 [Trinidad; pools]
  • Paracyclops rochai Karaytug & Boxshall, 1998 [Brazil; pools]
  • Paracyclops sitiseiensis Harada, 1931 [Formosa; fresh waters] *
  • Paracyclops smileyi Strayer, 1988 [USA; interstitial]
  • Paracyclops timmsi Kiefer, 1969 accepted as Australoeucyclops timmsi (Kiefer, 1969)
  • Paracyclops uenoi Ito, 1962 [Japan; cave waters]
  • Paracyclops vagus Lindberg, 1939 [India; fresh waters] **
  • Paracyclops waiariki lewis, 2010 [Nord Island; therm waters] (=Australoeucyclops waiariki (Lewis M.H., 1974)[PDF]
  • Paracyclops yeatmani Daggett & Davis, 1974 [USA; fresh waters] [PDF]
  • Paracyclops n.sp. Mercado-Salas &. Surez-Morales 2009 [Mexico]

    * syn. of P. affinis, according to Moncheko (1974)

    ** syn. of P. fimbriatus (Lindberg, 1958; Karaytug & Boxshall, 1998)


    OTHER SUBSPECIES OF PARAYCLOPS FIMBRIATUS

  • Paracyclops fimbriatus abnobensis Kiefer, 1929
  • Paracyclops fimbriatus andinus Lindberg, 1958
  • Paracyclops fimbriatus baicalensis Mazepova, 1962
  • Paracyclops fimbriatus bromeliarum Herbst, 1959
  • Paracyclops fimbriatus finitimus Kiefer, 1928
  • Paracyclops fimbriatus imminutus Kiefer, 1929
  • Paracyclops fimbriatus orientalis Alekseev, 1995

    UP DATE REVIEWS OF THE GENUS CAN BE FOUND IN :

    Karaytug & Boxshall (1998). The Paracyclops fimbriatus-complex (Copepoda, Cyclopoida): a revision. Zoosystema, 20 (4): 563-602.

    Karaytug & Boxshall (1998). Partial revison of Paracyclops Claus, 1893 (Copepoda, Cyclopoida, Cyclopidae) with description of four new species. Bull. nat. Hist. Mus. Lond. (Zool.), 64 (2): 111-205.



    ECTOCYCLOPS Brady, 1904


  • Ectocyclops affinis (Sars G.O., 1863)
  • Ectocyclops bradyi Mahoon & Zia, 1985
  • Ectocyclops bromelicola Kiefer, 1935 [Brazi, Venezuela?; bromeliads, springs]
  • Ectocyclops clausi Mahoon & Zia, 1985
  • Ectocyclops compactus (G.O. Sars, 1909) [Africa, Madagascar, south America; fresh waters]
  • Ectocyclops coperes (Gurney, 1928) [Tanganika; lakes]
  • Ectocyclops forbesi Mahoon & Zia, 1985
  • Ectocyclops ilariensis Onabamiro, 1952[Nigeria, fresh waters] *
  • Ectocyclops herbsti Dussart, 1984 [Venezuela, Guiana, Brazil, Argentina;ponds, lakes, rivers, springs] [PDF] [PDF2]
  • Ectocyclops hirsutus Kiefer, 1930 [Africa, Madagascar; lakes 3
  • Ectocyclops hirsutus euchaetus Kiefer, 1952 [Angola, fresh waters]
  • Ectocyclops hirsutus mixtus Dussart, 1974 [Ethiopie; fresh waters]
  • Ectocyclops medius Kiefer, 1930b
  • Ectocyclops ilariensis Onabamiro, 1952
  • Ectocyclops mozhae Baribwegure & Dumont, 2000 [Island of Soqotra, Indian Ocean; interstitial] (left) [PDF]
  • Ectocyclops phaleratus phaleratus (Koch, 1838) (syn. Paracyclops phaleratus (Koch, 1838) [cosmpolitan; fresh waters, lakes, marshes]
  • Ectocyclops phaleratus ilariensis Onabamiro, 1952 [Nigeria; fresh waters]
  • Ectocyclops phaleratus medius Ranga Reddy & Radhakrishna, 1985 [south India; lakes]
  • Ectocyclops phaleratus polyspinosus harada, 1931 [China; fresh waters]
  • Ectocyclops polyacanthus Harada, 1931
  • Ectocyclops polyspinosus Harada, 1931
  • Ectocyclops rahmi Lindberg, 1957 [Africa; lakes]
  • Ectocyclops rubescens Brady, 1904 [cosmpolitan, except Australia; fresh waters, ephemeral waters] [PDF] [PDF2]
  • Ectocyclops sarsi Mahoon & Zia, 1985
  • Ectocyclops strenzkei Herbst, 1959 [Brazil; bromeliads]
  • The following species, described by Mahon & Zia (1985) from Pakistan are questionable: E. Bradyi, E. clausi, E. forbesi, E. sarsi

    * questionable species



    THAUMASIOCYCLOPS Kiefer, 1930


  • Thaumasiocyclops insulanus Kiefer, 1930 [Indonesia; fresh waters]



    OCHRIDACYCLOPS Kiefer, 1937

  • Ochridacyclops arndi Kiefer, 1937 [Yugoslavia; lakes] (syn. Ochridacyclops arndti arndti Kiefer, 1937)
  • Ochridacyclops arndi f. prespensis Petkovski, 1954 [Yugoslavia; lakes]
  • Ochridacyclops brevicaudatus Shen & Tai, 1964 [China; lakes]
  • Ochridacyclops coreensis Lee, Leon & Yong Chang, 2004 [Korea; springs] (left) [PDF]
  • Ochridacyclops iriomotensis Ishida, 2002 [Japan; river puddle] [PDF]
  • Ochridacyclops kenyaensis Tomikawa et. al., 2005 [Kenya] [PDF]
  • Ochridacyclops nepalensis Tomikawa et. al., 2005 [Nepal] [PDF]
  • Ochridacyclops nipponensis Karaytug, Boxshall & Ishida, 1996 [Japan, streams] [PDF]


  • AUSTRALOEUCYCLOPS Karanovic, 2006 (subgenus of Eucyclops according Alekseev, 2019) [PDF]

    The eucyclopinid copepod genus Australoeucyclops Karanovic, 2006 currently contains six taxa: A. darwini Karanovic & Tang, 2009 from Western Australia; A. eucyclopoides (Kiefer, 1929) from Indonesia; A. linderi ( Lindberg, 1 948) from South Australia; A. timmsi (Kiefer, 1969) from New South Wales, Australia; A. waiariki ( Lewis, 1 974) from New Zealand; and A. karaytugi Karanovic, 2006 from Western Australia (see Karanovic 2006). However, Karanovic (2006) mentioned that another as yet undescribed member of this genus existed in the Margaret River region of Western Australia (from: Karanovic & Tang, 2009) [PDF]


  • Australoeucyclops darwini Karanovic & Tang, 2009 [Western Australia; ground waters] [PDF]
  • Australoeucyclops karaytugi Karanovic, 2006 [Australia; springs] (left) [PDF]
  • Australoeucyclops eucyclopoides (Kiefer, 1927) [Lombok; fresh waters]
  • Australoeucyclops linderi (Lindberg, 1948) [Australia; caves]
  • Australoeucyclops timmsi (Kiefer, 1969) [Australia, fresh waters]
  • Australoeucyclops waiariki (Lewis, 1974) [New Zealand; fresh waters] [PDF]


  • LINK TO THE COPEPOD WORLD AT SMITHSONIAN INSTITUTION


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