DIOSACCIDAE Sars, 1906

[now MIRACIIDAE Dana, 1846 (Willen, 2002)]


The Miracidae (ex Diosaccidae Sars, 1906) is a predominantly marine family, but containing several freshwater species, most of them in the genus Schizopera.

The family presently includes three subfamilies and 47 genera [DIOSACCINAE: Actopsyllus, Amonardia, Amphiascus, Amphiascopsis, Antiboreodiosaccus, Balucopsylla, Bulbamphiascus, Dactylopodamphiascopsis, Diosaccus, Diosaccopsis, Eoschizopera, Haloschizopera, Helmutkunzia, Ialysus, Metamphiascopsis, Miscegenus, Neomiscegenus, Paradiosaccus, Paramphiascella, Paramphiascoides, Paramphiascopsis, Pararobertsonia, Parialysus, Pholenota, Protopsammotopa, Psammotopa, Pseudamphiascopsis, Pseudodiosaccopsis, Pseudodiosaccus, Rhyncholagena, Robertgurneya, Robertsonia, Schizopera, Schizoperoides, Schizoperopsis, Sinamphiascus, Teissierella, Tydemanella, Typhlamphiascus; STENHELIINAE: Stenhelia, Pseudostenhelia, Cladorostrata, Onychostenhelia, Melima, Anisostenhelia, Beatricella, Hicksia; MIRACIINAE: Miracia, Oculosetella, Macrosetella, Distioculus ] with representatives for the most part living in sea waters, a few (Stenhelia, Schizopera, Eoschizopera, Schizoperoides, Cladorostrata, Paramphiascella, Amphiascus) including species living also in fresh or brackish waters, as well as in ground waters (Willen, 2000).

Key to genera in : Lang K. 1965. Copepoda Harpacticoidea from The Californan Pacific coast. K. svenska vetensk. Akad. Handl., 10 (2): 1-566



  • DIOSACCINAE Sars, 1906

  • STENHELIINAE Brady, 1880

  • MIRACIINAE Dana, 1846




    DIOSACCINAE Sars, 1906


    Actopsyllus Wells, 1967

    Amonardia Lang, 1944

    Amphiascoides Lotufo & Fleeger, 1995

    Amphiascopsis Gurney, 1927

    Amphiascus Sars, 1905

    Antiboreodiosaccus Lang, 1944

    Balucopsylla Rao, 1972

    Bulbamphiascus Lang, 1944

    Dactylopodamphiascopsis Lang, 1944

    Diosaccus Boeck, 1872

    Diosaccopsis Monard, 1928

    Distiocolus Boxshall, 1979

    Eoschizopera Wells & Rao, 1976

    Haloschizopera Lang, 1948

    Helmutkunzia Wells & Rao, 1976

    Ialysus Brian, 1927

    Macrosetella Sewell, 1940

    Metamphiascopsis Lang, 1944

    Miscegenus Wells, Hicks & Coull, 1982

    Miracia Boxshall, 1979

    Neomiscegenus Karanovic & Ranga Reddy, 2004

    Parialysus Nicholls, 1941

    Paramphiascella Lang, 1948

    Paramphiascoides Wells, 1967

    Paramphiascopsis Lang, 1944

    Pararobertsonia Lang, 1948

    Pholenota Vervoort, 1964

    Protopsammotopa Geddes, 1968

    Psammotopa Pennak, 1942

    Pseudamphiascopsis Lang, 1944

    Pseudodiosaccopsis Lang, 1944

    Rhyncolagena Lang, 1944

    Robertgurneya Lang, 1944

    Robertsonia Brady, 1880

    Schizopera Sars, 1905 s.l.

    Schizoperoides Por, 1968

    Schizoperopsis Apostolov, 1982

    Sinamphiascus Mu & Gee, 2000

    Teissierella Monard, 1935

    Typhlamphiascus Lang, 1944




    AMPHIASCUS Sars, 1905

  • Amphiascus amblyops Sars, 1911 [Norway; marine]
  • Amphiascus ampullifer (Humes, 1953) [north Europe; parasitic] *
  • Amphiascus angustipes Gurney, 1927 [Egypt, Italy, Bermuda, France, Sri Lanka; brackish waters]
  • Amphiascus brevis Sars, 1909 [Groenland; marine] *
  • Amphiascus congener Sars, 1909 [north Europe; ?]
  • Amphiascus caudaespinosus Brian, 1927 [Algerie; psammon]
  • Amphiascus demersus Nicholls, 1940 [St. Lawrence; sea waters]
  • Amphiascus dentiformis Coull, 1971 [Virgin Islands, meiobenthic]
  • Amphiascus discrepans Mielke, 1989 [Chile; marine benthos]
  • Amphiascus elegans Lang, 1948
  • Amphiascus elongatus Ito, 1972 [Japan, Antarctic (Livingston Island); interstitial, parasitic]
  • Amphiascus gauthieri Monard, 1936 [Algerie; psammon]
  • Amphiascus glacialis Lang, 1948
  • Amphiascus gracilis Lang, 1936 [Kerguelen; psammon]
  • Amphiascus graciloides Klie, 1950 [Ireland, Helgoland; marine]

  • Amphiascus hirtus Gurney, 1927 [Suez, Arabian Gulf, South Pacific, Brazil; marine]
  • Amphiascus ignotus Lang, 1948
  • Amphiascus longiremis Lang, 1948
  • Amphiascus longirostris Monard, 1928
  • Amphiascus lobatus Hicks, 1971 [new Zealand; coastal waters] (left) [PDF]
  • Amphiascus longarticulatus Marcus, 1974 [Ireland, Black sea; marine]
  • Amphiascus minutus (Claus, 1863) [Palaearctic, marine, psammon]
  • Amphiascus monodi Monard, 1952 [Africa, Senegal; coastal waters]*
  • Amphiascus mucronatus Lang, 1948
  • Amphiascus pacificus Sars, 1905 [Chatham Island; brackish waters, lagoons]
  • Amphiascus paracaudaespinosus Roe, 1958 [north Europe; coastal waters]
  • Amphiascus parvus Sars, 1906 [Europe, Bermuda, north America; marine, psammon]
  • Amphiascus perplexus (Thompson & A. Scott, 1903) [Sri Lanka, France?; marine]
  • Amphiascus polaris Sars, 1909 [Groenland;?]
  • Amphiascus profundus Becker & Schriever, 1979 [?]
  • Amphiascus propinquus Sars, 1906 [north Europe; marine]
  • Amphiascus sinuatus Sars, 1906 [north Europe, north America; sea waters]
  • Amphiascus tainui Hicks, 1989 [New Zealand; sea waters]
  • Amphiascus tenellus Sars, 1906 [Europe, Algerie; psammon]
  • Amphiascus tenuiremis (Brady & Robertson, 1880) [Europe; marine, psammon]
  • Amphiascus ultimus Monard, 1928 [France; marine] *
  • Amphiascus undosus Lang, 1965 [USA; coastal waters]
  • Amphiascus varians (Norman & Scott, 1905) [north Europe; marine, psammon, amongst algae]

    * questionable species


    PARAROBERTSONIA Lang, 1948


  • Pararobertsonia abyssi Lang, 1948
  • Pararobertsonia chesapeakensis Lang, 1948


    PSAMMOTOPA Pennak, 1942

  • Psammotopa biarticulata Mielke, 1990 [Panama; marine interstitial]
  • Psammotopa chappuisi Noodt, 1955 [France, Spain, north Africa; marine interstitial]
  • Psammotopa phyllosetosa (Noodt, 1952) [Europe; marine interstitial]
  • Psammotopa polyphylla Noodt, 1955 [France, Portugal; marine interstitial]
  • Psammotopa trisetosa Mielke, 1995 [Venezuela; marine interstitial]
  • Psammotopa vulgaris Pennak, 1942 [Europe, Israel, USA; marine interstitial]


    PROTOPSAMMOTOPA Geddes, 1968 [PDF]

  • Protopsammotopa norvegica Geddes, 1968 [Norway; marine interstitial]
  • Protopsammotopa wilsoni Wells, 1977 [USA; marine interstitial]
  • Protopsammotopa tipperi Bouck & Thistle, 2003 [Mexico; marine] [PDF]


    BALUCOPSYLLA Rao, 1972

  • Balucopsylla similis Rao, 1972 [India; marine interstitial]
  • Balucopsylla triarticulata Wells & Rao, 1987 [Andaman, Nicobar islands, Costa Rica; marine interstitial]


    NEOMISCEGENUS Karanovic & Ranga Reddy, 2004 [PDF]

  • Neomiscegenus indicus Karanovic & Ranga Reddy, 2004 [India; ground waters] [PDF]




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