DIOSACCIDAE Sars, 1906

[now MIRACIIDAE Dana, 1846 (Willen, 2002)]

DIOSACCINAE Sars,1906


TYPHLAMPHIASCUS Lang, 1944

  • Typhlamphiascus accraensis Lang, 1948
  • Typhlamphiascus blanchardi (Sars, 1906)
  • Typhlamphiascus brevicornis Lang, 1948
  • Typhlamphiascus bouligandi Soyer, 1972
  • Typhlamphiascus confusus (T. Scott, 1902) [north Europe, Israel, Red and Black seas; coastal waters]
  • Typhlamphiascus dentipes Lang, 1948
  • Typhlamphiascus drachi Soyer, 1964 [France; sea waters]
  • Typhlamphiascus gracilicaudatus (Thompson & A. Scott, 1903) [Ceylon, Red sea; coastal waters]
  • Typhlamphiascus gracilis Por, 1963 [north Koster; sea waters]
  • Typhlamphiascus lamellifer (Sars, 1911) [north Europe; sea waters]
  • Typhlamphiascus lamellifer capensis Kunz, 1975
  • Typhlamphiascus latifurca Por, 1968 [Dahlak Archipelago, Red sea; coastal waters]
  • Typhlamphiascus longifurcatus Rouch , 1962
  • Typhlamphiascus pectinifer Lang, 1965 [USA; marine]
  • Typhlamphiascus lutincola Soyer, 1963 [France; coastal waters]
  • Typhlamphiascus ovale Wells & Rao, 1987
  • Typhlamphiascus typhlops (Sars, 1906) [Europe; sea waters]
  • Typhlamphiascus typhloides Lang, 1948
  • Typhlamphiascus unisetosus Lang, 1965 [USA; marine]


    SCHIZOPERA Sars, 1905

    According to Mielke (1995) "more than 70 species belong to the taxon Schizopera, which shows at least two apomorphic features: the existence of only two spines on the outer margin of the distal segment of exopodites P2-P4 and the presence of a hyaline spine on the distal segment of exopodite P3 (male)". The former is shared with other genera, the latter is an important autapomorphy for the genus Schizopera.

    Apostolov (1982) divided the genus Schizopera into three genera: Schizopera (with two subgenera: Schizopera s.str.and Neoschizopera); Eoschizopera Apostolov, 1982 (with two subgenera: Eoschizopera s.str. and Praeoschizopera); and Schizoperopsis Apostolov, 1982 (with two subgenera: Schizoperopsis s. str. and Psammoschizoperopsis). Mielke (1992) rejected these groupings, even if favoured them as a "tool to facilitate identification of the species". The same opinion is shared by Bodin (1997).


  • Schizopera (Schizopera) aegyptica Kiefer, 1978 [Egypt; lakes]
  • Schizopera (Schizopera) akatovae Borutzky, 1953 [Caspian sea]
  • Schizopera (Schizopera) anomala Coull, 1971 [USA; coastal waters]
  • Schizopera (Schizopera) aralensis Borutzky, 1971 [Aral sea; Uzbekistan] [PDF]
  • Schizopera (Schizopera) austindownsi Karanovic, 2004 [W-Australia; ground waters]
  • Schizopera (Schizopera) baltica Lang, 1965 [Europe; brackish waters]
  • Schizopera (Schizopera) bozici Lang, 1965 [Reunion; coastal waters]
  • Schizopera (Schizopera) borutzkii Monchenko, 1967 [Black sea]
  • Schizopera (Schizopera) bradyi Soyer, 1975 [Kerguelen; mesopsammon]
  • Schizopera (Schizopera) brusinae Petkovski, 1954 [Bulgaria, Black Sea; brackish waters]
  • Schizopera (Schizopera) californica Lang, 1965 [USA; coastal waters]
  • Schizopera (Schizopera) carolinensis Coull, 1971 [USA; coastal waters]
  • Schizopera (Schizopera) cicolanii Galassi & Pesce, 1988 [south Italy; brackish waters] [PDF]
  • Schizopera (Schizopera) clandestina clandestina (Klie, 1924) [Europe, Africa, China, New zealand; brackish waters]
  • Schizopera (Schizopera) clandestina brevicauda Kiefer, 1965
  • Schizopera (Schizopera) compacta Lint, 1922 [Europe;brackish waters]
  • Schizopera (Schizopera) consimilis Sars, 1909 [Africa; lakes]
  • Schizopera (Schizopera) costaricana Karanovic, 2004 [W-Australia; ground waters]
  • Schizopera (Schizopera) depotspringsi Karanovic, 2004 [W-Australia; ground waters]
  • Schizopera (Schizopera) dimentmani Por, 1993 [Israel; shore springs]
  • Schizopera (Schizopera) fimbriata Sars, 1909 [Africa; lakes]
  • Schizopera (Schizopera) grimalschii Jakubisiak, 1938 [Romania; marine?]
  • Schizopera (Schizopera) haitiana Kiefer, 1934 [Haiti, Venezuela; lakes]
  • Schizopera (Schizopera) hawaiiensis Kunz, 1955 [Hawai; lagoons]
  • Schizopera (Schizopera) inopinata Sars, 1909 [Africa; Tanganika lake]
  • Schizopera (Schizopera) issykkulica Manuilova, 1966 [URSS; lakes]
  • Schizopera (Schizopera) jugurtha jugurtha (Blanchard & Richard, 1891) [Europe, Africa; fresh and brackish waters]
  • Schizopera (Schizopera) jugurtha stephanidesi (Pesta, 1938) [Europe; brackish waters]
  • Schizopera (Schizopera) jundeei Karanovic, 2004 [W-Australia; ground waters]
  • Schizopera (Schizopera) knabeni Lang, 1965 [USA; coastal waters]
  • Schizopera (Schizopera) kunzi Apostolov, 1967 [Black sea]
  • Schizopera (Schizopera) lagrecai Pesce & Galassi, 1987 [Italy, Sicily; ground waters] [PDF]
  • Schizopera (Schizopera) langi Petkovski, 1954 [Bulgaria; brackish waters]
  • Schizopera (Schizopera) lindae Apostolov & Pesce, 1984 [south Italy; ground waters] [PDF]
  • Schizopera (Schizopera) longicauda Sars, 1905 [Egypt; brackish waters]
  • Schizopera (Schizopera) longifurcata (Chappuis, 1955) [Africa; lakes]
  • Schizopera (Schizopera) longirostris (Daday, 1901) [Thailand, New Guinea; brackish waters]
  • Schizopera (Schizopera) marlieri Rouch & Chappuis, 1960 [Africa; Tanganika lake]
  • Schizopera (Schizopera) minuta Noodt, 1955 [France, coastal waters]
  • Schizopera (Schizopera) minuticornis Sars, 1909 [Africa; Tanganika lake]
  • Schizopera (Schizopera) nana Noodt, 1955 [France, coastal waters]
  • Schizopera (Schizopera) neglecta Akatova, 1935 [Black Sea, Asia; lakes]
  • Schizopera (Schizopera) noodti Rouch, 1962 [south America; coastal waters]
  • Schizopera (Schizopera) oldouei Karanovic, 2004 [W-Australia; ground waters]
  • Schizopera (Schizopera) osana Mielke, 1995 [Costa Rica; coastal waters]
  • Schizopera (Schizopera) paradoxa (Daday, 1903) [Asia; lakes]
  • Schizopera (Schizopera) parvula Noodt, 1955 [France, coastal waters]
  • Schizopera (Schizopera) petkovskii Apostolov, 1971 [Bulgaria; brackish waters]
  • Schizopera (Schizopera) pontica Chappuis & Serban, 1953 [Romania, Bulgaria; coastal waters]
  • Schizopera (Schizopera) pori Jemenez Alvarez, 1988 [Brazil; coastal rivers] [PDF]
  • Schizopera (Schizopera) pseudojugurtha Borutzky, 1972 [URSS: lakes]
  • Schizopera (Schizopera) reducta Borutzky, 1971 [Aral sea]
  • Schizopera (Schizopera) rotundipes Gurney, 1926 [Africa; Tanganika lake]
  • Schizopera (Schizopera) scalaris Sars, 1909 [Africa; Tanganika lake]
  • Schizopera (Schizopera) spinulosa Sars, 1909 [Africa; Tanganika lake]
  • Schizopera (Schizopera) subterranea Lang, 1948 [Europe; ground waters]
  • Schizopera (Schizopera) taricheana Por, 1968 [Israel; lakes]
  • Schizopera (Schizopera) tobae tobae Chappuis, 1931 [Indonesie, Cuba, Bulgaria; lakes]
  • Schizopera (Schizopera) tobae cubana Petkovski, 1973 [Cuba, Yucatan; caves, cenotes]
  • Schizopera (Schizopera) tobae wolterecki Brehm & Chappuis, 1935 [Indonesie, brackish waters]
  • Schizopera (Schizopera) triacantha Kiefer, 1934 [Haiti; brackish waters]
  • Schizopera (Schizopera) ungulata Sars, 1909 [Africa; Tanganika lake]
  • Schizopera (Schizopera) uramurdahi Karanovic, 2004 [W-Australia; ground waters]
  • Schizopera (Schizopera) validior Sars, 1909 [Africa; Tanganika lake]
  • Schizopera (Schizopera) variseta Bozic, 1964 [Reunion; coastal waters]
  • Schizopera (Schizopera) vicina Herbst, 1960 [Perù, Brazil; brackish coastal lagoons]

  • Schizopera (Neoschizopera) arconae Arlt, 1983 [Baltic sea]
  • Schizopera (Neoschizopera) chaetosa Petkovski, 1954 [Black sea]
  • Schizopera (Neoschizopera) giselae Jimenez Alvarez, 1988 [Brazil; brackish coastal waters] [PDF]
  • Schizopera (Neoschizopera) lacusamari Por & Marcus, 1972 [Suez canal]
  • Schizopera (Neoschizopera) meridionalis meridionalis (Petkovski, 1954) [Bulgaria; brackish waters]
  • Schizopera (Neoschizopera) meridionalis listensis Mielke, 1975 [north Europe; sea waters]
  • Schizopera (Neoschizopera) monardi Petkovski, 1955 [Yugoslavie; brackish waters]
  • Schizopera (Neoschizopera) ornata Noodt & Purasjoki, 1953 [Europe; brackish waters]
  • Schizopera (Neoschizopera) pratensis Noodt, 1958 [Germany; sea waters]
  • Schizopera (Neoschizopera) spinifer Wells & Rao, 1987 [Andaman, Nicobar; sea waters]


    SUBGENUS NOT DEFINED

  • Schizopera inornata Noodt, 1955
  • Schizopera roberiverensis Karanovic, 2006 [Australia; bores]
  • Schizopera rybnikovi Chestopood & Korney, 2006 [Caspian Sea]
  • Schizopera setulosa Mirabdullayev & Ginatullina, 2007 [Uzbekistan; lake] [PDF]
  • Schizopera spinulosa Mirabdullayev & Ginatullina, 2007 [Uzbekistan; lake] *[PDF]
  • Schizopera weelumurra Karanovic, 2006 [Australia; wells]

    *Preoccupied by Schizopera (Schizopera) spinulosa Sars, 1909


    EOSCHIZOPERA Wells & Rao, 1976

  • Eoschizopera (Eoschizopera) chiloensis Mielke, 1992 [Chile; sea waters]
  • Eoschizopera (Eoschizopera) nicoyana Mielke, 1995 [Costa Rica; sea waters]
  • Eoschizopera (Eoschizopera) reducta Wells & Rao, 1976 [Azores?]
  • Eoschizopera (Eoschizopera) syltensis (Mielke, 1973) [north Europe; hyporheic]

  • Eoschizopera (Praeoschizopera) crassispinata (Chappuis, 1954) [Madagascar; psammon]
  • Eoschizopera (Praeoschizopera) elatensis (Kahan & Bar-El, 1982
  • Eoschizopera (Praeoschizopera) gliglici Petkovski, 1957 [Yugoslavie, Bulgaria; brackish waters]
  • Eoschizopera (Praeoschizopera) indica (Rao & Ganapati, 1969) [India; coastal sea waters]
  • Eoschizopera (Praeoschizopera) soyeri (Kunz, 1983)

    Incertae sedis: E. (P.) marlieri (Rouch & Chappusi, 1960); S. issykkulica Mauyilova, 1966


    SCHIZOPEROPSIS Apostolov, 1982

  • Schizoperopsis (Schizoperopsis) arenicola (Chappuis & Serban, 1953) [Romania; coastal waters]
  • Schizoperopsis (Schizoperopsis) nichollsi (Soyer, 1975) [Kerguelen; mesopsammon]
  • Schizoperopsis (Schizoperopsis) varnensis Apostolov, 1967 [Black sea]

  • Schizoperopsis (Psammoschizoperopsis) gauldi (Chappuis & Rouch, 1961) [Africa, Ghana, Brazil ;marine interstitial]


    SCHIZOPEROIDES Por, 1968

  • Schizoperoides expeditionis Por, 1968 [Dahlak Archipelago, Red sea]


    HALOSCHIZOPERA Lang, 1948

    Gee & Fleeger (1990), considering the number of inner setae on the exopod 3 of legs 3-4, grouped the species of the genus known at that time into four categories:

    1 - Exopod 3 of leg 3-4 with 1 and 2 inner setae respectivey (H. pygmaea, H. mathoi, H. marmarae, H. ruthorum, H. bulbifer)
    2 - Exopod 3 of leg 3-4 with no setae and 2 inner setae respectivey (H. abyssi, H. lima)
    3 - Exopod 3 of leg 3-4 with 1 seta (H. noodti , H. aegyptica)
    4 - Exopod 3 of leg 3-4 with no setae and 1 inner seta respectivey (H. exigua, H. pauciseta, H. phyllura, H. conspicua, H. minima, H. bathyalis, H. apprisea)

  • Haloschizopera abyssi Becker, 1974 [Europe?; sea waters]
  • Haloschizopera aegyptica Noodt, 1964 [Red Sea]
  • Haloschizopera apprisea Gee & Fleeger, 1990 [Alaska; sea waters]
  • Haloschizopera bathyalis Schriever, 1984 [north Atlantic ocean, Iceland; marine]
  • Haloschizopera bulbifera (Sars, 1911) [Norway; psammon]
  • Haloschizopera clotensis Moore & Reilly, 1993 [?]
  • Haloschizopera conspicua Por, 1964 [Skagerak; psammon]
  • Haloschizopera exigua (Sars, 1906) [north Europe, Italy; psammon]
  • Haloschizopera junodi Monard, 1935 [north Europe, Black Sea] *
  • Haloschizopera latisetifera Marinov, 1973 [Atlantic ocean]
  • Haloschizopera lima Becker, 1974 [Europe?; sea waters]
  • Haloschizopera lionensis Moore & Reilly, 1993 [?]
  • Haloschizopera marmarae Noodt, 1955 [Marmara Sea]
  • Haloschizopera mathoi (Monard, 1935) [Tunisia; sea waters]
  • Haloschizopera minima Por, 1964 [Skagerak; psammon]
  • Haloschizopera noodti Bodin, 1968 [France; psammon]
  • Haloschizopera nuditerga Moore & Reilly, 1993 [?]
  • Haloschizopera pauciseta Por, 1959/1964 [Black sea]
  • Haloschizopera phyllura Noodt, 1964 [Red Sea]
  • Haloschizopera pygmaea (Norman & T. Scott, 1905) [north Europe; marine, muddy sand]
  • Haloschizopera ruthorum Por, 1967 [Red Sea]
  • Haloschizopera tenuipes Noodt, 1964 [Red Sea]

    * questionable species



    PAGE UPDATED APRIL 11. 2008


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