 CANTHOCAMPTIDAE Sars, 1906, emend. Monard, 1927, Lang, 1948 (syn.HETEROPSYLLIDAE Kornev & Chertoprud, 2008)
Cletocamptinae Gomes & Rivera, 2022 
AMPHIBIPERITA Fiers & Rutledge, 1990
Amphibiperita neotropica (Jakobi, 1956) [USA, Brazil; brackish coastal waters]
AUSTRALOCAMPTUS Karanovic, 2004
Australocamptus hamondi Karanovic, 2004 [W-Australia; ground waters]
Australocamptus similis Karanovic, 2004 [W-Australia; ground waters]
Australocamptus diversus Karanovic, 2004 [W-Australia; ground waters]
BOREOLIMELLA Huys & Thistle, 1989
Boreolimella nympha (Por, 1964) [Skagerak; muds]
Boreolimella dubia (Wells, 1965) [Scotland; muds]
PUSILLARGILLUS Huys & Thistle, 1989
Pusillargillus nixe Huys & Thistle, 1989 [Israel, ?]
PERUCAMPTUS Huys & Thistle, 1989
Perucamptus rapiens Becker, Noodt & Schriever, 1979 [north America; ?]
DAHLAKOCAMPTUS Huys, 2009 (syn. Dahlakia Por, 1986)
Dahlakocamptus xenuus (Por, 1968) [Red Sea]
HANIKRAIA Huys, 2009 (syn. Poria Lang, 1965)
Hanikraia derketo (Por, 1964) [levantine and Pontic; sea waters]
MESOPSYLLUS Por, 1960
Mesopsyllus atargatis Por, 1960 [Schwarzen Sea]
Mesopsyllus curvisetus (Kornev & Chertoprud, 2008)
Mesopsyllus dimorphus Mu & Huys, 2017 [China]
Mesopsyllus glacialis Nokinov & Sharafutdigen, 2021 [Artic]
Mesopsyllus secundus (Wells, 1965) [Scotland; meiobenthos]
Mesopsyllus spiniferus Mu & Huys, 2017 [China]
PAREPACTOPHANES Kunz, 1935
Parepactophanes dubia Noodt, 1958 [Canarie islands; coastal waters] (syn.Taurocletodes dubius (Noodt, 1958)
syn. of
Taurocletodes dubius (Noodt, 1958)
Parepactophanes gallicus (Kunz, 1975)
Parepactophanes minuta Kunz, 1935
Parepactophanes tumenae (Karaytug & Huys, 2004) accepted as Taurocletodes tumenae Karaytug & Huys, 2004
HEMIMESOCHRA Sars, 1920
Hemimesochra atargatis (Por, 1960) accepted as Mesopsyllus atargatis Por, 1960
Hemimesochra clavularis Sars, 1920 [Norway; psammon]
Hemimesochra derketo Por, 1964 = Poria derketo (Por, 1964)
Hemimesochra dubia Wells, 1965 accepted as Boreolimella dubia (Wells, 1965)
Hemimesochra nympha Por, 1964 accepted as Boreolimella nympha (Por, 1964)
Hemimesochra nixe Por, 1964 accepted as Pusillargillus nixe (Por, 1964)
Hemimesochra rapiens Becker, Noodt & Schriever, 1979 accepted as Perucamptus rapiens (Becker, Noodt & Schriever, 1979)
Hemimesochra secunda Wells, 1965 accepted as Mesopsyllus secundus (Wells, 1965)
Hemimesochra trisetosa Coull, 1973 [North Carolina, USA] (=Carolinicola trisetosa (Coull, 1973)
NANNOMESOCHRA Gurney, 1932 (syn. Archeolourinia Corgosinho & Schizas, 2013, according to Alper et al., 2023)
Nannomesochra arupinensis (Brian, 1925) [Europe, Africa, north America, Australia; marine, wetlands]
Nannomesochra erythraiensis Alper et al., 2023 [Turkey]
Nannomesochra gebekumensis Alper et al., 2023 [Turkey]
Nannomesochra gizizi Alper et al., 2023 [Turkey]
Nannomesochra parvula Gurney, 1927 (syn. Nannomesochra arupinensis (Brian, 1925)
Nannomesochra shermani Alper et al., 2023 [Turkey]
Nannomesochra zavodniki Petkovski & Apostolov, 1974
HETEROPSYLLUS T. Scott, 1894 (syn. Cletomesochra Sars G.O., 1920)
"The genus Heteropsyllus T. Scott, 1894 can be
referred to as the mass genera Canthocamptidae s. lat.
This genus, which is quite complex for taxonomy, was
initially attributed to Cletodidae T. Scott, 1904 [Lang,
1948] and later transferred to Canthocamptidae [Por,
1986]. Kornev & Chertoprud [2008] separated the genus into a monotypic family Heteropsyllidae Kornev et
Chertoprud, 2008. Following that, some authors still
indicated this genus as part of the Canthocamptidae
family [George, 2013; Garlitska et al., 2019], including the recently described species Heteropsyllus spongiophilus Novikov et Sharafutdinova, 2021. The authors indicated the similarity of Heteropsyllus in some
characteristics to Hemimesochrinae; however, they do
not indicate the exact position [Novikov, Sharafutdinova 2021]" (from Novikov et al., 2021
The genus should be separated into the family Heteropsyllidae, according to Kornev & Chertoprud [2008].
Heteropsyllus confluens (Soyer, 1975) [France]
Heteropsyllus coulli Kornev & Chertoprud, 2008
Heteropsyllus coreanus Nam & Lee, 2006 [Korea; interstitial]
Heteropsyllus curticaudatus (Scott, 1894) [UK; marine]
Heteropsyllus celticus Nam & Lee, 2006 [England; interstitial]
Heteropsyllus dimorphus Por, 1959 (syn. Heteropsyllus major (Sars G.O., 1920)
Heteropsyllus exiguus (Sars, 1911) [Norway]
Heteropsyllus major (Sars, 1920) [Norway; psammon]
Heteropsyllus masculus Kunz, 1971 [Germany]
Heteropsyllus meridionalis (Soyer, 1975) [France]
Heteropsyllus minutus Wells, 1965 accepted as Bathycamptus minutus (Wells, 1965)
Heteropsyllus nanus (Sars, 1920) [Norway, Sweden]
Heteropsyllus nunni (Coull,1975) [USA]
Heteropsyllus pseudonunni Coull & Palmer, 1980 [USA]
Heteropsyllus rostratus rostratus (Sars, 1920) [north Europe; psammon]
Heteropsyllus rostratus similis Smirnov, 1964 [Arctic Ocean]
Heteropsyllus serratus Schriever. 1983
Heteropsyllus spiridonovi Novikov et al., 2021 [Laptev Sea]
Heteropsyllus spongiophilus Nokinov & Sharafutdigen, 2021 [Artic]
| Key to the species of the genus Heteropsyllus (modified from Coull and Palmer 1980) |
PSAMMOCAMPTUS Mielke, 1975
Psammocamptus axi Mielke, 1975 [Sylt Island, Germany; psammon]
Psammocamptus eckmani (Huys & Thistle, 1989) (syn. Bathycamptus eckmani Huys & Thistle, 1989
Psammocamptus galapagoensis Mielke, 1997 [Galapagos; marine interstitial]
Psammocamptus minutus (Wells , 1965) (syn. Bathycamptus minutus (Wells, 1965)
Canthocamptidae
incertae sedis
TAUROCLETODES Kunz, 1975 (Copepoda, Canthocamptidae incertae sedis according to Karaytug & Huys 2004)
Taurocletodes dubius (Noodt, 1958) [Canarie islands; coastal waters]
Taurocletodes gallicus (Noodt, 1958) (syn. Parepactophanes gallicus (Kunz, 1975)
Taurocletodes tumenae Karaytung & Huys, 2004 [Turkey; sandy beaches]
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