EURYTEIDAE Monchenko 1975


EURYTE Philippi 1843 (syn. Ancheuryte Herbst, 1989;Thorellia Boeck, 1865)


The genus Euryte presently includes ten named species and subspecies, worldwide distributed in shallow and deeper waters; some species have been found living in sand interstices or associated with corals and seaweed.


  • Euryte brevicauda Sewell, 1949 [Indian Ocean]
  • Euryte curticornis Sars, 1913 [East Atlantic Ocean]
  • Euryte grata Herbst, 1989 [West Indies, Puerto Rico; marine coastal waters]
  • Euryte jaumei Karanovic, 2014 [Spain; marine]
  • Euryte koreana Karaman, 2014 [Korea; marine]
  • Euryte longicauda Philippi 1843 emend. Giesbrecht, 1900 [European coasts, Suez; sea, caves]
  • Euryte longicauda minor T. Scott, 1905 [U.K., Norway; sea] [PDF]
  • Euryte longiseta Grandori, 1925
  • Euryte notabilis (Herbst, 1989)
  • Euryte propinqua Brady, 1910
  • Euryte pseudorobusta Vervoort 1964 [Pacific Ocean]
  • Euryte robusta Giesbrecht, 1900 [Cosmopolitan; sea waters]
  • Euryte sewelli Vervoort, 1964 [Indian Ocean]
  • Euryte similis Scott, 1912 [East Atlantic Ocean]
  • Euryte verecunda Humes, 1992 [Panama; coral associated] (=Coraleuryte verecunda (Humes, 1992) [PDF]

  • ANCHEURYTE Herbst, 1989 (syn. Euryte Philippi, 1843 )

  • Ancheuryte notabilis Herbst, 1989 [West Indies, Puerto Rico; marine coastal waters] (= Euryte notabilis (Herbst, 1989)


    CORALEURYTE Karanovic, 2014

  • Coraleuryte bellatula (Humes, 1991)
  • Coraleuryte verecunda (Humes, 1992)


    FRATIIDAE Ho, Conradi & López-González, 1998

    FRATIA Ho, Conradi & Lopez-Gonzalez, 1998

  • Fratia gaditana Ho, Conradi & López-González, 1998


    CUCUMARICOLIDAE Bouligand & Delamare-Deboutteville, 1959

    CUCUMARICOLA Paterson, 1958

  • Cucumaricola curvatus Avdeev, 1977
  • Cucumaricola notabilis Paterson, 1958


    BOTRYLLOPHILIDAE Sars G.O., 1921 (syn.Haplostomidae Sars G.O., 1921: Haplostominae Chatton & Hatton, 1924)

    BOTRYLLOPHILUS Hesse, 1864 (syn. Blakeanus Wilson C.B., 1921; Kossmechthrus Della Valle, 1883 )

  • Botryllophilus abbotti Ooishi & Illg, 1989
  • Botryllophilus angustus Kim I.H. & Boxshall, 2021
  • Botryllophilus antarcticus Kim I.H. & Boxshall, 2021
  • Botryllophilus aspinosus Schellenberg, 1922
  • Botryllophilus bamfieldensis Ooishi, 2000
  • Botryllophilus banyulensis Brément, 1909 [PDF]
  • Botryllophilus bergensis Schellenberg, 1921
  • Botryllophilus bermudensis Kim I.H. & Boxshall, 2021
  • Botryllophilus brevipes Brément, 1909 (syn. Botryllophilus sarsi Ooishi, 2002
  • Botryllophilus conicus Conradi, Lopez-Gonzalez & Garcia-Gomez, 1994 [PDF]
  • Botryllophilus coniorhynchus Kim I.H. & Boxshall, 2021
  • Botryllophilus constellatus Hesse, 1866
  • Botryllophilus corniger (Wilson, C.B., 1921) (syn.Botryllophilus norvegicus Schellenberg, 1921)
  • Botryllophilus curtipes Kim I.H. & Boxshall, 2021
  • Botryllophilus dentirostris Kim I.H. & Boxshall, 2021
  • Botryllophilus distinctus Ooishi, 2012
  • Botryllophilus groenlandicus (Hansen, 1923) (syn. Botryllophilus norvegicus Schellenberg, 1921)
  • Botryllophilus guadeloupensis Kim I.H. & Boxshall, 2021
  • Botryllophilus inaequipes Hansen, 1923
  • Botryllophilus koreensis Seo & Lee, 1995
  • Botryllophilus kozloffi Ooishi, 2014
  • Botryllophilus longicaudatus Kim I.H. & Boxshall, 2021
  • Botryllophilus longipes Kim I.H. & Boxshall, 2021
  • Botryllophilus macropus Canu, 1892
  • Botryllophilus millari Ooishi, 2014
  • Botryllophilus neapolitanus Ooishi, 2006
  • Botryllophilus norvegicus Schellenberg, 1921
  • Botryllophilus nudisetatus Kim I.H. & Boxshall, 2021
  • Botryllophilus pentachaetus Kim I.H. & Boxshall, 2021
  • Botryllophilus randalli Stock, 1970
  • Botryllophilus ruber Hesse, 1864
  • Botryllophilus sarsi Ooishi, 2002
  • Botryllophilus spinulosus Ooishi, 2012
  • Botryllophilus stenurosus Kim I.H. & Boxshall, 2021
  • Botryllophilus symmetricus Ooishi, 2014
  • Botryllophilus tuberculatus Kim I.H. & Boxshall, 2021

    HAPLOSACCUS Chatton & Harant, 1924

  • Haplosaccus ardius Kim I.H. & Boxshall, 2021
  • Haplosaccus elongatus Ooishi & Illg, 1977
  • Haplosaccus sacculus (Chatton & Brément, 1910)

    HAPLOSTOMA Chatton & Harant, 1924
    (syn. Aphelostoma Brement, 1909; Aplostoma Canu, 1886); Ceuthopodus Hesse, 1866; Cryptopodus Hesse, 1865; Tranestoma Wilson C.B., 1924)

  • Haplostoma affinis (Scott T., 1906)
  • Haplostoma alba (Hesse, 1868)
  • Haplostoma albicatum Ooishi & Illg, 1977
  • Haplostoma amarouci (Blake, 1929) (syn. Haplostomides amarouci (Blake, 1929)
  • Haplostoma ambiguum Ooishi & Illg, 1977
  • Haplostoma banyulensis (Brément, 1909)
  • Haplostoma bispinosum Ooishi, 2009 [PDF]
  • Haplostoma brevicauda (Canu, 1886)
  • Haplostoma canui Chatton & Harant, 1924
  • Haplostoma dentatum Ooishi & Illg, 1977
  • Haplostoma depressum Kim I.H. & Boxshall, 2021
  • Haplostoma dudleyae Ooishi, 1998
  • Haplostoma elegans Ooishi & Illg, 1977
  • Haplostoma elongatum Ooishi, 2009 [PDF]
  • Haplostoma eruca (Norman, 1869)
  • Haplostoma fusiforme Kim I.H. & Boxshall, 2021
  • Haplostoma gibberum (Schellenberg, 1922)
  • Haplostoma gibbera (Schellenberg, 1922) (syn. Haplostoma gibberum (Schellenberg, 1922)
  • Haplostoma gracile Kim I.H. & Boxshall, 2021
  • Haplostoma hibernica (Scott T. & Scott A., 1895) (syn. Haplostomides hibernicus (Scott T. & Scott A., 1895)
  • Haplostoma humesi Ooishi, 1995
  • Haplostoma junctum Ooishi, 2009 [PDF]
  • Haplostoma mammiferum Kim I.H. & Boxshall, 2021
  • Haplostoma megallanica (Chatton & Brément, 1910) (syn. Haplostomella magellanica (Chatton & Brément, 1910)
  • Haplostoma kimi Seo & Lee, 2001
  • Haplostoma laticaudatum Kim I.H. & Boxshall, 2021
  • Haplostoma madagascarensis Ooishi, 2009 [PDF]
  • Haplostoma manadoense Kim I.H. & Boxshall, 2021
  • Haplostoma minutum Ooishi & Illg, 1977
  • Haplostoma mizoulei Monniot C., 1962
  • Haplostoma pingue Kim I.H. & Boxshall, 2021
  • Haplostoma pygmaeum Kim I.H. & Boxshall, 2021
  • Haplostoma rectangulatum Kim I.H. & Boxshall, 2021
  • Haplostoma saccula (Chatton & Brément, 1910) (syn. Haplosaccus sacculus (Chatton & Brément, 1910)
  • Haplostoma setiferum Ooishi & Illg, 1977
  • Haplostoma simplex Kim I.H. & Boxshall, 2021
  • Haplostoma sycozoae Salfi, 1926 (syn. Haplostomella sycozoae (Salfi, 1926)
  • Haplostoma symbioticum Kim I.H. & Boxshall, 2021
  • Haplostoma vanuatuense Kim I.H. & Boxshall, 2021


    HAPLOSTOMELLA Chatton & Harant, 1924 (syn. Rhabdomorpha Fukui, 1965)

  • Haplostomella australiensis Gotto, 1970
  • Haplostomella bilobata Kim I.H. & Boxshall, 2021
  • Haplostomella binodosa Kim I.H. & Boxshall, 2021
  • Haplostomella biseta Kim I.H. & Boxshall, 2021
  • Haplostomella borealis Marchenkov & Boxshall, 2003
  • Haplostomella crassa Kim I.H. & Boxshall, 2021
  • Haplostomella distincta Ooishi & Illg, 1977
  • Haplostomella dubia Ooishi & Illg, 1977
  • Haplostomella halocynthiae (Fukui, 1965)
  • Haplostomella magellanica (Chatton & Brément, 1910)
  • Haplostomella malacocera Chatton & Harant, 1924
  • Haplostomella multinodosa Kim I.H. & Boxshall, 2021
  • Haplostomella oceanica Ooishi & Illg, 1977
  • Haplostomella ooishiae Kim I.H. & Boxshall, 2021
  • Haplostomella reducta Ooishi & Illg, 1977
  • Haplostomella sycozoae (Salfi, 1926)
  • Haplostomella tuberculata Chatton & Harant, 1924
  • Haplostomella uniseriata Kim I.H. & Boxshall, 2021

    HAPLOSTOMIDES Chatton & Harant, 1924

  • Haplostomides amarouci (Blake, 1929)
  • Haplostomides bellus Ooishi & Illg, 1977
  • Haplostomides bermudensis Kim I.H. & Boxshall, 2021
  • Haplostomides brementi Chatton & Harant, 1924
  • Haplostomides gottoi Ooishi, 2008
  • Haplostomides hibernicus (Scott T. & Scott A., 1895)
  • Haplostomides hawaiiensis Ooishi, 1994
  • Haplostomides hibernicus (Scott T. & Scott A., 1895)
  • Haplostomides luteolus Ooishi & Illg, 1977
  • Haplostomides otagoensis Ooishi, 2001[PDF]
  • Haplostomides partitus Kim I.H. & Boxshall, 2021
  • Haplostomides sanamyani Marchenkov & Boxshall, 2003
  • Haplostomides scotti Chatton & Harant, 1924
  • Haplostomides similis Kim I.H. & Boxshall, 2021

    PAULILLGIA Monniot C., 1982

  • Paulillgia polycarpae Monniot C., 1982

    SCHIZOPROCTUS Aurivillius, 1885 (syn. Pteropygus Sars G.O., 1921)

  • Schizoproctus bisetatus Kim I.H. & Boxshall, 2021
  • Schizoproctus fijiensis Kim I.H. & Boxshall, 2021
  • Schizoproctus frigidus Kim I.H. & Boxshall, 2021
  • Schizoproctus fusiformis Kim I.H. & Boxshall, 2021
  • Schizoproctus inflatus Aurivillius, 1885
  • Schizoproctus magnus Kim I.H. & Boxshall, 2021
  • Schizoproctus mollis Kim I.H. & Boxshall, 2021
  • Schizoproctus oligomerus Kim I.H. & Boxshall, 2021
  • Schizoproctus pinguis Kim I.H. & Boxshall, 2021
  • Schizoproctus tripartitus Kim I.H. & Boxshall, 2021
  • Schizoproctus vestitus (Sars G.O., 1921)

    Blakeanus Wilson C.B., 1921 (= Botryllophilus Hesse, 1864; Ceuthopodus Hesse, 1866 (= Haplostoma Chatton & Harant, 1924; Cryptopodus Hesse, 1865 (= Haplostoma Chatton & Harant, 1924) Haplostominae Chatton & Hatton, 1924 (syn. Botryllophilidae Sars G.O., 1921; Kossmechthrus Della Valle, 1883 (syn. Botryllophilus Hesse, 1864; Pteropygus Sars G.O., 1921 (syn.Schizoproctus Aurivillius, 1885; Rhabdomorpha Fukui, 1965(syn. Haplostomella Chatton & Harant, 1924; Tranestoma Wilson C.B., 1924 (syn. Haplostoma Chatton & Harant, 1924.


    BUPRORIDAE Thorell, 1859

    BUPRORIDES Kim I.H. & Boxshall, 2021

  • Buprorides trilobatus Kim I.H. & Boxshall, 2021

    BUPRORUS Thorell, 1859

  • Buprorus caudatus Illg & Dudley, 1980
  • Buprorus loveni Thorell, 1859


    ENTEROGNATHIDAE Illg & Dudley, 1980

    ENTEROGNATHUS Giesbrecht, 1900

  • Enterognathus comatulae Giesbrecht, 1900
  • Enterognathus inabai Ohtsuka, Shimomura & Kitazawa, 2012
  • Enterognathus lateripes Stock, 1966

    GOMPHOPODARION Humes, 1974

  • Gomphopodarion byssoicum Humes, 1974

    PARENTEROGNATHUS Ohtsuka, Kitazawa & Boxshall, 2010

  • Parenterognathus troglodytes Ohtsuka, Kitazawa & Boxshall, 2010

    ZANCLOPUS Calman, 1908

  • Zanclopus antarcticus Gravier, 1912
  • Zanclopus cephalodisci Calman, 1908


    ENTEROPSIDAE Thorell, 1859

    ENTEROCOLA van Beneden, 1860 (syn. Biocryptus Hesse, 1865; Intercola Hesse, 1862)

  • Enterocola adnatus Ooishi, 2014
  • Enterocola africanus López-González, Conradi & Garcia-Gómez, 1993
  • Enterocola australis Kim I.H. & Boxshall, 2021
  • Enterocola beaumonti Scott T. & Scott A., 1895 (syn. Haplostomides beaumonti (Scott T. & Scott A., 1895)
  • Enterocola betencourti Canu, 1891 (syn. Enterocola fulgens van Beneden, 1860)
  • Enterocola bilamellata Sars G.O., 1921
  • Enterocola brementi Illg & Dudley, 1980
  • Enterocola clavelinae Chatton & Harant, 1924
  • Enterocola coniculus Ooishi, 2014
  • Enterocola curticaudus Kim I.H. & Boxshall, 2021
  • Enterocola daudinae Chatton & Harant, 1924 (syn. Enterocola clavelinae Chatton & Harant, 1924
  • Enterocola dicaudatus Marchenkov & Boxshall, 2005 [PDF]
  • Enterocola eruca Norman, 1869 (syn. Haplostoma eruca (Norman, 1869)
  • Enterocola fertilis Illg & Dudley, 1980
  • Enterocola fulgens van Beneden, 1860
  • Enterocola gottoi Conradi, Lopez-Gonzalez & Garcia-Gomez, 1992
  • Enterocola hessei Chatton & Harant, 1924
  • nterocola hibernica Scott T. & Scott A., 1895 (syn.Tranestoma hibernica (Scott T. & Scott A., 1895)
  • Enterocola ianthina Illg & Dudley, 1980
  • Enterocola laticeps Illg & Dudley, 1980
  • Enterocola mabulensis Kim I.H. & Boxshall, 2021
  • Enterocola mammifera Chatton & Harant, 1922
  • Enterocola megalova Gotto, 1962
  • Enterocola monnioti Marchenkov & Boxshall, 2005 [PDF]
  • Enterocola neocaledonicus Kim I.H. & Boxshall, 2021
  • Enterocola nodulosus Kim I.H. & Boxshall, 2021
  • Enterocola oligosetosus Kim I.H. & Boxshall, 2021
  • Enterocola ooishiae O'Reilly, 2008 [PDF]
  • Enterocola parapterophorus Marchenkov & Boxshall, 2005 [PDF]
  • Enterocola parvus Kim I.H. & Boxshall, 2021
  • Enterocola petiti Guille, 1964
  • Enterocola precaria Illg & Dudley, 1980
  • Enterocola pterophora Chatton & Brément, 1909
  • Enterocola quadrisetus Kim I.H. & Boxshall, 2021
  • Enterocola robustus Kim I.H. & Boxshall, 2021
  • Enterocola sedentarius Kim I.H. & Boxshall, 2021
  • Enterocola seticaudus Kim I.H. & Boxshall, 2021
  • Enterocola setifera Hansen, 1923 (syn. Enterocola setiferus Hansen, 1923)
  • Enterocola sulcatus Kim I.H. & Boxshall, 2021
  • Enterocola sydnii Chatton & Harant, 1924
  • Enterocola tuberculatus Kim I.H. & Boxshall, 2021
  • Enterocola unisetosus Kim I.H. & Boxshall, 2021

    ENTEROCOLIDES Chatton & Harant, 1922

  • Enterocolides ecaudatus Chatton & Harant, 1922
  • Enterocolides elongatus Kim I.H. & Boxshall, 2021
  • Enterocolides pacificus Kim I.H. & Boxshall, 2021

    ENTEROPSIS Aurivillius, 1885 (syn. Haligryps Aurivillius, 1885; Ichnograde Hesse, 1864; Plantipode Hesse, 1864;Synthetys Canu, 1886)

  • Enteropsis abbotti Illg & Dudley, 1980
  • Enteropsis aculeatus (Aurivillius, 1885) (syn.Enteropsis sphinx Aurivillius, 1885)
  • Enteropsis arcticus Marchenkov, 1994
  • Enteropsis capitulatus Illg & Dudley, 1980
  • Enteropsis chattoni Monniot C., 1961
  • Enteropsis elongata Kim I.H. & Boxshall, 2021
  • Enteropsis fusiformis Ooishi, 2009
  • Enteropsis georgianus Schellenberg, 1922
  • Enteropsis hispida Kim I.H. & Boxshall, 2021
  • Enteropsis minor Illg & Dudley, 1980
  • Enteropsis nudus Kim I.H. & Moon, 2011
  • Enteropsis obtusa Kim I.H. & Boxshall, 2021
  • Enteropsis onychophorus Schellenberg, 1922
  • Enteropsis roscoffensis Chatton & Brément, 1909
  • Enteropsis ruber (Hesse, 1864)
  • Enteropsis sphinx Aurivillius, 1885
  • Enteropsis superbus Illg & Dudley, 1980
  • Enteropsis tasmanica Kim I.H. & Boxshall, 2021
  • Enteropsis teres (Aurivillius, 1885) (syn. Enteropsis sphinx Aurivillius, 1885)
  • Enteropsis tromsoensis Kim I.H. & Boxshall, 2021
  • Enteropsis vararensis Scott T., 1901 (syn. Mychophilus roseus Hesse, 1866)

    LEQUERREA Chatton & Harant, 1924

  • Lequerrea canui Illg & Dudley, 1980z
  • Lequerrea perezi Chatton & Harant, 1924

    MONNIOTICOPIA Kim I.H. & Boxshall, 2021

  • Monnioticopa firma Kim I.H. & Boxshall, 2021
  • Monnioticopa manadoensis Kim I.H. & Boxshall, 2021
  • Monnioticopa minuta Kim I.H. & Boxshall, 2021
  • Monnioticopa planicoxa Kim I.H. & Boxshall, 2021
  • Monnioticopa prima Kim I.H. & Boxshall, 2021

    MYCHOPHILUS Hesse, 1865 (syn Adranesius Hesse, 1865)

  • Mychophilus capillatus Kim I.H. & Moon, 2011
  • Mychophilus curvatus Chatton & Brément, 1909 (syn. Mychophilus roseus Hesse, 1866
  • Mychophilus elatus (Hesse, 1869)
  • Mychophilus fallax Stock, 1967
  • Mychophilus hesperius Kim I.H. & Boxshall, 2021
  • Mychophilus notialis Kim I.H. & Boxshall, 2021
  • Mychophilus pachygaster Hesse, 1866 (syn.Mychophilus roseus Hesse, 1866
  • Mychophilus palmatus López-González & Conradi, 1996 [PDF]
  • Mychophilus roseus Hesse, 1866
  • Mychophilus rosovula Gray, 1933 (syn. Mychophilus roseus Hesse, 1866
  • Mychophilus ruber (Hesse, 1865) (syn. Botryllophilus ruber Hesse, 1864
  • Mychophilus vararensis (Scott T., 1901) (syn. Mychophilus roseus Hesse, 1866

    PERIBOIA Kim I.H. & Boxshall, 2021

  • Periboia tahitiensis Kim I.H. & Boxshall, 2021

    Biocryptus Hesse, 1865 accepted as Enterocola van Beneden, 1860 Subfamily Enterocolinae Chatton & Hatton, 1924 =Enteropsidae Thorell, 1859 ;Haligryps Aurivillius, 1885 = Enteropsis Aurivillius, 1885; Ichnograde Hesse, 1864 =Enteropsis Aurivillius, 1885; Intercola Hesse, 1862 = Enterocola van Beneden, 1860; Plantipode Hesse, 1864= Enteropsis Aurivillius, 1885; Synthetys Canu, 1886= Enteropsis Aurivillius, 1885,




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