 CHAPPUISIIDAE Chappuis, 1940 CHAPPUISIUS Kiefer, 1938
Chappuisius inopinus Kiefer, 1938 [Europe; ground waters]
Chappuisius singeri Chappuis, 1939 [Europe: mesopsammon]
CLETODIDAE Scott, 1905 (sensu Por, 1986)
Por (1986) in his review of the family Cletodidae transferred some genera of this family to a new sub-family of Canthocamptidae: Hemimesochrinae, as well as provisionally placed the genus Cletocamptus in the "Canthocamptidae incertae sedis". Later on, Gee (1999), sharing the Por's (1986) opinion, provisionally included the genus Cletocamptus in the family Canthocamptidae incertae sedis, suggestings as well that this genus "must be removed to a new family which will brobably prove to be a sister family to the Cletodidae".
More recently, Mielke (2000) confirmed that "The situation of the genus Cletocamptus is complex and in some respects inconsistent..... This uncertainty is due to the conjecture that the two families, Canthocamptidae and Cletodidae, appear to be poly- or para-phyletic assemblages".
The genus Pontocletodes Apostolov, 1980 was not defined by Por (1986) and it is provisionally assigned to Cletodidae. Bodin (1997) reported the same genus as "incertae sedis", adding some new genera to the family Cletodidae.
Pending a definitive review of the above families, the genus Cletocamptus is herein provisonally placed in the
Cletodidae sensu Scott, 1904; Por, 1986, which presently should include the following other genera: Acrenhydrosoma, Cletodes, Enhydrosoma, Enhydrosomella, Kollerua, Limnocletodes, Stylicletodes [Monocletodes Lang], Barbaracletodes, Australonannopus, Scintis, Strongylacron, Schizacron, Monocletodes, Intercletodes, Dyacrenhydrosoma, Paracrenhydrosoma , Triathrix, Sphingothrix, Neoacrenhydrosoma (Por, 1986, Fiers, 1987; Bodin, 1997; Gee, 1998,1999) and Spinapecruris Gee, 2001.
CLETODES Brady, 1872
Cletodes brucei Lang, 1948
Cletodes carthaginiensis Monard, 1935 [north Africa, Tunisia; marine]
Cletodes confusum Gomez, 2000 [USA; lagoons] [PDF]
Cletodes dentatus Wells & Rao, 1987 [Andaman, Nicobar; coastal waters]
Cletodes dissimilis Willey, 1935 [Bermuda; ?]
Cletodes dorae Por, 1979 [Red sea]
Cletodes endopodita (Schriever, 1984) [north Atlantic ocean]
Cletodes hartmannae Lang, 1965 [USA; sea waters]
Cletodes latirostris Drzycimiski, 1967 [Norway; sea waters]
Cletodes limicola limicola Brady, 1872 [north Europe, Algerie, Indian ocean; sea waters]
Cletodes limicola tunisiensis Monard, 1935 [Tunisia; marine]
Cletodes longicaudatus (Boeck, 1872) [Palaearctic: sea waters]
Cletodes longifurca Lang, 1948 [north Europe; sea waters]
Cletodes macrura Fiers, 1991 [USA; coastal waters]
Cletodes millerorum Hamond, 1973 [Australia; sea waters]
Cletodes pseudodissimilis Coull, 1971 [USA; meiobenthic] [PDF]
Cletodes pseudodissimilis oris Gomez, 2000 [USA; lagoons]
Cletodes pusillus Sars, 1920 [north Europe; sea waters]
Cletodes reductus Moore, 1978 [France; sea waters]
Cletodes reyssi Soyer, 1964 [France; coastal waters]
Cletodes setosus Marinov & Apostolov, 1985 [spanish Sahara; coastal waters]
Cletodes smirnovi Bodin, 1970 [France; sea intertidal]
Cletodes spinulipes Por, 1967 [Elat, Red sea; marine]
Cletodes tenuipes T. Scott, 1896 [north Europe; sea waters]
Cletodes tuberculatus Fiers, 1991 [USA; coastal waters]
Cletodes yotabis Por, 1967 [Elat, Red sea; marine ]
ENHYDROSOMA Boeck, 1872
Enhydrosoma baruchi Coull, 1975 [USA; estuarine waters]
Enhydrosoma brevipodum Gomez, 2004 [?: ?]
Enhydrosoma buchholtzi (Boeck, 1872) [north Europe, north America; marine]
Enhydrosoma caeni Raibaut, 1965 [Thau basin; sea waters]
Enhydrosoma cananeiae jakobi, 1955 [Brazil; brackish costal waters] *
Enhydrosoma casoae Gómez, 2003 [PDF]
Enhydrosoma curticauda Boeck, 1872 [Europe, Italy; sea waters]
Enhydrosoma curvirostris (T. Scott, 1894) [Europe; psammon] **
Enhydrosoma franklini Thistle, 1980
Enhydrosoma gariene Gurney, 1930 [England; brackish waters]
Enhydrosoma gerlachi Jakobi, 1955 [Brazil; brackish costal waters] *
Enhydrosoma guaratubae Jakobi, 1955 [Brazil; brackish costal waters] *
Enhydrosoma herrerai Bell & Kern, 1983 [USA; estuarine waters]
Enhydrosoma hopkinsi Lang, 1965 [USA; sea waters]
Enhydrosoma intermedia Chislenko, 1978 [Japan; sea waters]
Enhydrosoma ivetteae Jakobi, 1955 [Brazil; brackish costal waters] *
Enhydrosoma kuehnemanni Pallares, 1970
Enhydrosoma lacunae Jakubisiak, 1933 [Cuba; lagoons] [PDF]
Enhydrosoma latipes (A. Scott, 1909) [Africa; sea waters, plankton]
Enhydrosoma latipes (A. Scott, 1909) [Africa; sea waters, plankton]
Enhydrosoma littorale Wells, 1967 [Africa, Mozambico; sea waters]
Enhydrosoma longicauda Marinov & Apostolov, 1983 [Atlantic ocean]
Enhydrosoma longifurcatum Sars, 1909 [Europe, Italy; psammon]
Enhydrosoma mangroviae Jakobi, 1955 [Brazil; brackish coastal waters] *
Enhydrosoma micrurum Monard, 1928 [France; marine]
Enhydrosoma migoti Monard, 1926 [France; marine]
Enhydrosoma minimum Jakobi, 1955 [Brazil; brackish costal waters] *
Enhydrosoma nicobaricum Sewell, 1940 [USA?; sea waters]
Enhydrosoma parapropinquum Gómez, 2003 [PDF]
Enhydrosoma pectinatum Wells & Rao, 1987 [Andaman, Nicobar islands; coastal waters]
Enhydrosoma pericoense Mielke, 1990 [Panama; coastal waters]
Enhydrosoma pontica Lang, 1948
Enhydrosoma propinquum (Brady, 1880) [Europe, Italy, Algerie; coastal waters]
Enhydrosoma rosae Fiers, 1996 [nom. nov. (Fiers, 1996) for E. propinquum]
Enhydrosoma sarsi (T. Scott, 1904) [north Europe; coastal waters]
Enhydrosoma solitarum Gomez, 2003 [PDF]
Enhydrosoma sordidum Monard, 1926 [European Atlantic coasts, Black and Mediterranean Seas, Brazil; coastal waters]
Enhydrosoma stylicaudatum Willey, 1935 [Bermuda, ?]
Enhydrosoma tunisense Monard, 1935 [Africa, Tunisia; marine]
Enhydrosoma uniarticulatum Borutzky, 1928 [URSS; brackish waters]
Enhydrosoma variabile Wells, Hicks & Coull, 1982 [New Zealand; estuarine waters]
Enhydrosoma wellsi Bodin, 1968 [France; sea waters]
* species inquirenda (Lang, 1965; Por et al., 1984)
** recently moved to the genus Spinapecruris Gee, 2001 (Gee, 2001)
CLETOCAMPTUS Schmankewitch, 1875
The systematic status of the genus Cletocamptus is still controversial. Several authors (Por, 1968; Bodin, 1997; Mielke, 2000, 2001; Gomez et al., 2007) include it in Canthocamptidae incertae sedis sensu Por, 1986. Pending a definitive review of the above family, the genus Cletocamptus is herein provisonally placed in the
Cletodidae sensu Scott, 1904.
At present the valid species within Cletocamptus are C. retrogressus Schmankewitsch, 1875,
C. confluens (Schmeil, 1894), C. albuquerquensis (Herrick, 1894), C. trichotus Kiefer, 1929,
C. feei (Shen, 1956), C. affinis Kiefer, 1957, C. gravihiatus (Shen and Sung, 1963), C.
helobius Fleeger, 1980, C. merbokensis Gee, 1999, C. axi Mielke, 2000, C. schmidti Mielke,
2000, C. deborahdexterae Gomez, Fleeger, Rocha-Olivares and Foltz, 2004, C. stimpsoni
Gomez, Fleeger, Rocha-Olivares and Foltz, 2004, C. gabrieli Loffler, 1961, C. sinaloensis Gomez, Fleeger, Rocha-
Olivares and Foltz, 2004, C. bicolor (Wilson, 1932), C. xenuus Por, 1968 , C. kummieri (Delachaux,1917), C. stimpsoni Gomez, Fleeger, Rocha Oliovarez & Faltz, 2003, C. fourchensis Gomez, Fleeger, Rocha-Olivares and Foltz,
2004, C. cecsurirensis Gomez, Scheihing & La Barca, 2007. C. mongolicus (Sterba, 1967), C. levis Gomez, 2005, C. nudus Gomez, 2005 Gomez et al. (2004) considered as species inquirendae some species within the genus
and they also considered as doubtful some records of C. deitersi. Additional doubtful
records of C. deitersi are those of Brehm (1936, 1965), Chappuis (1936), Ringuelet (1958a,
1958b, 1960, 1962), Ringuelet et al. (1965), Oliveira and Miranda (1971), Apostolov
(1984), Ruber et al. (1994), and Loftus and Reid (2000).
Cletocamptus affinis Kiefer, 1957 [China; brackish waters]
Cletocamptus albuquerquensis (Herrick, 1894) [Americas; fresh, brackish waters]
Cletocamptus axi Mielke, 2000 [Galapagos; lagoons] [PDF] [PDF2]
Cletocamptus bicolor (Wilson, 1932) [America; fresh, brackish waters]
Cletocamptus cecsurirensis Gomez, Scheihing & La Barca, 2007 [Chile; fresh waters] [PDF]
Cletocamptus confluens (Schmeil, 1894) [cosmopolitan, except America; fresh, brackish waters, wetlands] [PDF] *
Cletocamptus deborahdexterae Gomez, Fleeger, Rocha Oliovarez & Faltz, 2003 [USA; fresh waters] (syn. C. deitersi partim) [PDF]
Cletocamptus deitersi deitersi (Richard, 1897) [Cuba, Yucatan, south America, China, Australia, Israel; Ethiopie; brackish waters, wetlands, cenotes]
Cletocamptus deitersi ecuadorianus Loffler, 1964 [Ecuador, brackish waters]
Cletocamptus feei (Shen, 1956) [China; brackish waters]
Cletocamptus fourchensis Gomez, Fleeger, Rocha Oliovarez & Faltz, 2003 [USA; fresh waters] [PDF]
Cletocamptus gabrieli Loffler, 1961 [Iran; brackish waters]
Cletocamptus gravihiatus (Shen & Sung, 1963) [China; brackish waters]
Cletocamptus helobius Fleeger, 1980 [USA; brackish waters]
Cletocamptus kummieri (Delachaux,1917) [south America; marine]
Cletocamptus levis Gomez, 2005 [Brazil; marine] [PDF]
Cletocamptus merbokensis Gee, 1999 [Malaysia; mangrove forest] [PDF]
Cletocamptus mongolicus (Sterba, 1967) [Mongolia]
Cletocamptus nudus Gomez, 2005 [Brazil; marine] [PDF]
Cletocamptus retrogressus Schmankevitch, 1875 [Europe, Africa, Asia; fresh, brackish waters] [PDF]
Cletocamptus schmidti Mielke, 2000 [Galapagos; lagoons] [PDF]
Cletocamptus sinaloensis Gomes et al., 2004 [Mexico, Brazil; fresh waters] [PDF]
Cletocamptus stimpsoni Gomez, Fleeger, Rocha Oliovarez & Faltz, 2003 [USA; fresh waters] [PDF]
Cletocamptus trichotus Kiefer, 1929 [south Africa; fresh, brackish waters]
Cletocamptus xenuus Por, 1968 [Dahlak Archipelago, Red Sea; brackish waters]
* Cletocamptus confluens, according to its variability and widespread occurrence, could be considered an "assemblage of barely discernible species" (Mielke, 2000). For the same reason, the subspecies Cletocamptus confluens cyrenaica Brehm, 1908 and Cletocamptus confluens meridionalis Kiefer, 1929 are questionable.
 Map of the distribution of Cletocamptus confluens (after Mielke, 2000 modif.)
The species Cletocamptus deitersi and Cletocamptus retrogressus are geographically widspread, and morphologically and ecologically variable thus implying the existence of morphologically indistinguishable sibling species (Mielke, 2001).
| Sincere thanks are due to the colleague S. Gomez for sending PDF files on Cletocamptus |
PONTOCLETODES Apotolov, 1980
Pontocletodesponticus Apostolov, 1980 [Black sea; psammon]
NANNOPODELLA Monard, 1928
Nannopodella denisi Monard, 1928
KOLLERUA Gee, 1994
Kollerua birsteini (Borutzky, 1971) [Aral Sea]
Kollerua breviarticulatum (Shen & Tai, 1964) [China; fresh waters]
Kollerua longum Shen & Tai, 1979 [China; fresh waters]
Kollerua radhakrishnai (Ranga Reddy, 1979) [India; lakes]
Kollerua uniarticulatum (Borutzky, 1928) [URSS; brackish waters]
STRONGYLACRON Gee & Huys, 1996 [PDF]
Strongylacron buchholzi (Boeck, 1872) [north Europe, sea waters] [PDF]
SCHIZACRON Gee & Huys, 1996 [PDF]
Schizacron barnishi (Wells, 1967) [Africa, Mozambico; coastal waters] [PDF]
Schizacron bifurcarostratum (Shen & Tai, 1965) [China; rivers] [PDF]
Schizacron intermedius Gee & Huys, 1996 [Thailand; mangrove] [PDF]
Schizacron vervoorti (Fiers, 1987) [India; brackish waters] [PDF]
ACRENHYDROSOMA Lang, 1944
The genera Acrenhydrosoma Lang, 1944, Dyacrenhydrosoma
Gee, 1999, Paracrenhydrosoma Gee, 1999 and
Neoacrenhydrosoma Gee and Mu, 2000 constitute a well
defined lineage (Acrenhydrosoma-lineage) within
the Cletodidae.
(see Bodin 1997 and Gee. 1999). The strongly modified endopodal
lobe of the fifth legs in both sexes, the position of the
exopods in the fith legs when present, the armature of
the female sixth legs, separate these four genera from
all other known cletodids (from: Kotwicki & Fiers, 2005).
Acrenhydrosoma karlingi Lang, 1965 [USA; sea waters]
Acrenhydrosoma maccalli Schizas & Shirley, 1994 [Alaska; sea waters] [PDF]
Acrenhydrosoma perplexa (T. Scott, 1899) [Europe; sand sediments]
KEY TO SPECIES OF ACRENHYDROSOMA [PDF]
DYACRENHYDROSOMA Gee, 1999 [PDF]
Dyacrenhydrosoma breviseta Gee, 1999 [S.E.Asia; mangrove forest]
PARACRENHYDROSOMA Gee, 1999 [PDF]
Paracrenhydrosoma maccalli (Schizas & Shirley, 1994) [Alaska; mudflat intertidal]
Paracrenhydrosoma oceaniae Kotwicki & Fiers, 2005 [Kongsfjorden (Svalbard)] [PDF]
Paracrenhydrosoma normani Gee, 1999 [Norway; sea waters] [PDF]
NEOACRENHYDROSOMA Gee & Mu, 2000 [PDF]
Neoacrenhydrosoma zhangi Gee & Mu, 2000 [China; muddy sand] [PDF]
SPINAPECRURIS Gee, 2001
Spinapecruris curvirostris (T. Scott, 1894) [North-West Europe; marine muddy sediment]
| RELEVANT LITERATURE | | NEXT PAGE |
CRISTACOXIDAE Huys, 1990
The family Crystacoxidae Huys, 1990 presently includes four genera living in marine psammon (Noodtorthopsyllus Lang, 1965; Cristacoxa, Huys, 1990; Laophontisochra George, 2002) and ground (Cubanocleta Petkovski, 1977) waters.
NOODTORTHOPSYLLUS Lang, 1965
Noodtorthopsyllus psammophilus (Noodt, 1955) [Canarie islands; psammon]
CUBANOCLETA Petkovski, 1977
Cubanocleta noodti Petkovski, 1977 [Cuba, cave waters]
CRISTACOXA Huys, 1990
Cristacoxa petkovskii Huys, 1990 [West Indies; psammon]
LAOPHONTISOCHRA George, 2002
Laophontisochra maryamae K. H. George, 2002 [Chile; psammon]
| PAGE UPDATEDJUNE 30. 2008 |
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