LAOPHONTOPSIDAE Huys & Willems, 1989


LAOPHONTOPSIS Sars, 1908 (syn. Cleta Claus, 1863)

  • Laophontopsis borealis Huys & Willems, 1989 [Europe; north Africa, marine]
  • Laophontopsis brevirostris (Claus, 1863) (= Paralaophonte brevirostris (Claus, 1863)
  • Laophontopsis forcipata (Claus, 1866)
  • Laophontopsis lamellifera (Claus, 1863) [Europe; north Africa, marine]
  • Laophontopsis lilljeborgia (Krichagin, 1877)
  • Laophontopsis minuticornis (Buchholz, 1874)
  • Laophontopsis monardi Huys & Willems, 1989 [Europe; north Africa, marine]
  • Laophontopsis parvula (Claus, 1866)
  • Laophontopsis secunda (Sewell, 1924) (syn, Telodocus secundus (Sewell, 1924)
  • Laophontopsis serrata (Claus, 1863) (syn. Laophonte serrata (Claus, 1863)
  • Laophontopsis setigera (Krichagin, 1877)
  • Laophontopsis similis (Krichagin, 1873) (syn. Heterolaophonte similis fortior (Krichagin, 1873)
  • Laophontopsis thalestris (Krichagin, 1877)
  • Laophontopsis uncinata (Czerniavski, 1868) (syn. Heterolaophonte uncinata (Czerniavski, 1868)


    ACULEOPSIS Huys & Willems, 1989

  • Aculeopsis longisetosa Huys & Willems, 1989


    TELODOCUS Huys & Willems, 1989

  • Telodocus secundus Huys & Willems, 1989




    THOMPSONULIDAE Lang, 1944 sensu Huys & Gee, 1990


    Huys & Gee (1990) raised to family rank the subfamily Thompsonulinae Lang, 1944, formerly included in the family Tachidiidae (Lang, 1948).

    The family presently includes two genera, viz. Thompsonula T. Scott, 1905 and Caribbula Huys & Gee, 1990.


    THOMPSONULA T. Scott, 1905 (syn. Echinocornus Wilson C.B., 1932; Rathbunula Wilson C.B., 1932)

  • Thompsonula agilis (Wilson C.B., 1932)
  • Thompsonula curticaudata (C.B. Wilson, 1932) [USA; marine sand]
  • Thompsonula hyaenae (I.C. Thompson, 1889) [Europe, USA, Africa; marine]
  • Thompsonula hyaenae elongata Gee, 1988 (syn. Caribbula elongata (Gee, 1988)
  • Thompsonula pectinata (Wilson C.B., 1932)


    CARIBBULA Huys & Gee, 1990

  • Caribbula elongata (Gee, 1988) [Mexico; marine coastal sand]
  • Caribbula fleegeri Huys & Gee, 1990 [USA; marine, continental shelf]



    RELEVANT LITERATURE


    Huys R. & J.M. Gee. 1990. A revision of Thompsonulidae Lang, 1944 (Copepoda: Harpacticoida). Zool. J. Linnean Soc., 99: 1-49.





    ZOSIMEIDAE Seifried, 2003

    "The family Zosimeidae, currently composed of the genera Zosime Boeck, 1873, Peresime Dinet, 1974, Heterozosime Kim &Lee. 2021, Acritozosime Kim & Lee, 2021 and Pseudozosime T. Scott, 1912, was erected by Seifried (2003) based on unique apomorphies in the mouth parts, P1 endopod, and P5 (Seifried 2003; Huys & Clark 2009). These genera can easily be distinguished from each other by the segmentation of the endopods of P2-P4 (Wells 2007; Koller & George 2011). Although the family is small, presently accommodating only 19 species/subspecies and mainly from sublittoral marine habitats (Wells 2007; Koller & George 2011), many unnamed species have been reported in the literature and are awaiting description (Carey & Montagna 1982; Mu et al. 2002; Seifried 2003; Chertoprud et al. 2009; Huys & Clark 2009). Zosime is the most speciose genus in the family, currently comprising 16 species/subspecies and assuming a wide geographical distribution (Koller & George 2011). Recently, Koller & George (2011) noted that some species display derived features in the antennule, fifth leg, and caudal ramus, and suggested that a phylogenetic study based on the type material of most species is necessary to confirm or reject the monophyletic status of the genus" (accordin to to Huys & Clark, 2016)

    ZOSIME Boeck, 1873

    "Based on morphological comparison of female character states, the genus Zosime can be subdivided into three provisional groups, anneae-, incrassata-, and typica-group. These groups are distinguished from each other by differences in the antennule, antennary exopod, mandibular rami, armature of swimming legs, fifth leg, and caudal rami" (Kim et al., 2016)

  • Zosime anneae Koller & George, 2011 [Atlantic Ocean]
  • Zosime atlanticola Bodin, 1968 (?)
  • Zosime bathybia Bodin, 1968 (?)
  • Zosime bathyalis Por, 1967
  • Zosime bergensis Drzycimski, 1968
  • Zosime carsteni Pointner, 2017
  • Zosime changi Kim & Lee, 2019
  • Zosime comata Kim & Lee, 2019
  • Zosime destituta Kim et al., 2016 [Korea]
  • Zosime eliasi Pointner, 2017
  • Zosime erythraea Por, 1967
  • Zosime fusiformis Brady, 1880 (syn. Jonesiella fusiformis (Brady & Robertson, 1876)
  • Zosime gisleni Lang, 1948
  • Zosime gymnokosmosa Kim & Lee, 2019
  • Zosime incrassata incrassata Sars, 1910
  • Zosime incrassata bathybia Bodin, 1968
  • Zosime major Sarts, 1919
  • Zosime mediterranea Lang, 1948
  • Zosime montagnai Yeom et al., 2022 [Mexico; marine]
  • Zosime mexicana Gomez & Rivera, 2023 [USA]
  • Zosime pacifica Fiers, 1991
  • Zosime paramajor Bodin, 1968
  • Zosime paratypica Becker & Schriever, 1979
  • Zosime reducta Becker & Schriever, 1979 (syn. Peresime reducta (Becker & Schriever, 1979)
  • Zosime reyssi Dinet, 1974
  • Zosime spinulosa Brady & Robertson, 1876 (syn, Danielssenia typica Boeck, 1873)
  • Zosime thistlei Yeom et al., 2022 [Mexico; marine]
  • Zosime tunnelli Yeom et al., 2022 [Mexico; marine]
  • Zosime typica Boeck, 1873
  • Zosime valida Sars, 1919
  • Zosime xiximi Gomez & Rivera, 2023 [USA]

    SPECIES INCERTAE SEDIS OR INQUIRENDAE

  • Zomine rubra Thompson, 1888 (nomen nudum)


    HETEROZOSIME Kim & Lee, 2021 [Indian Ocean]

  • Heterozosime klemie Gomez & Rivera, 2023 [USA]
  • Heterozosime tenuis Kim & Lee, 2021 [Indian Ocean]

    ACRITOZOSIME Kim & Lee, 2021 [Indian Ocean]

  • Acritozosime sigsbeeiensis Gomez & Rivera, 2023 [USA]
  • Acritozosime spinesco Kim & Lee, 2021 [Indian Ocean]

    PSEUDOZOSIME T. Scott, 1912

  • Pseudozosime brownei T. Scott, 1912

    PERESIME Dinet, 1974

  • Peresime abyssalis Dinet, 1974
  • Peresime pryocta Gomez & Rivera, 2023 [USA]
  • Peresime pryorae Yeom et al., 2022 [Mexico; marine]
  • Peresime reducta (Becker & Schriever, 1979)




    PORCELLIDIIDAE Boeck, 1865


    BREVIFRONS Harris V.A., 1994

  • Brevifrons faviolatum Harris V.A., 1994

    CEREUDORSUM Harris V.A., 2014

  • Cereudorsum verrucosum Harris V.A., 2014

    CLAVIGOFERA Harris V.A. & Iwasaki, 1996

  • Clavigofera clavigera (Pesta, 1935)
  • Clavigofera echinophila (Humes & Gelerman, 1962)
  • Clavigofera laurencia (Hicks, 1982)
  • Clavigofera pacifica Harris V.A. & Iwasaki, 1996
  • Clavigofera ulva (Hicks, 1982)

    CLUNIA Harris V.A., 2014

  • Clunia cocosensis Harris V.A., 2014

    DILATATIOCAUDA Harris V.A., 2002

  • Dilatatiocauda bipartita (Kim I.H. & H.S. Kim, 1997)
  • Dilatatiocauda dilatata (Hicks, 1971)
  • Dilatatiocauda medialis Harris V.A., 2002
  • Dilatatiocauda multidenticulata Harris V.A., 2002
  • Dilatatiocauda plana (Tiemann, 1977)
  • Dilatatiocauda retroseta Harris, 2002 [Australia]
  • Dilatatiocauda tristanensis (Wiborg, 1964)

    KEY TO SPECIES OF DILATATIOCAUDA

    GEDDESIA Harris V.A., 2014

  • Geddesia quadrata Harris V.A., 2014
  • Geddesia trisetosa (Geddes, 1968)

    KENSAKIA Harris V.A. & Iwasaki, 1997

  • Kensakia acuta (Kim I.H. & Kim H.S., 1997)
  • Kensakia acuticaudata (Thompson I.C. & Scott A., 1903)
  • Kensakia aiiroa Harris V.A. & Iwasaki, 1997
  • Kensakia australis Harris V.A., 2014
  • Kensakia parva Harris V.A. & Iwasaki, 2009
  • Kensakia shimodensis Harris V.A. & Iwasaki, 2009

    KIOLOARIA Harris V.A., 1994

  • Kioloaria bipunctata (Harris V.A., 2014)
  • Kioloaria brevicaudata (Thompson I.C. & Scott A., 1903)
  • Kioloaria cumula (Harris V.A., 2014)
  • Kioloaria edenensis (Harris V.A., 2014)
  • Kioloaria geddesi (Harris V.A., 2014)
  • Kioloaria iwasakii (Harris V.A., 2014)
  • Kioloaria jejuensis Huys, 2016
  • Kioloaria paguri (Ho, 1986)
  • Kioloaria quinquelineata (Harris V.A. & Robertson H.M., 1994)
  • Kioloaria rufolineata (Harris V.A. & Robertson, 1994)
  • Kioloaria sesquimaculata Harris V.A., 1994
  • Kioloaria similis (Kim S.H. & Kim W., 1996)
  • Kioloaria tapui (Hicks & Webber, 1983)

    Key to species of Kioloaria Harris, 1994


    KUSHIA Harris V.A. & Iwasaki, 1996

  • Kushia gamoi Harris V.A. & Iwasaki, 1996
  • Kushia igaguria Harris V.A. & Iwasaki, 1996
  • Kushia spathoides Harris V.A., 2014
  • Kushia zosteraphila Harris V.A. & Iwasaki, 1996

    MUCROROSTRUM Harris V.A. & Iwasaki, 1997

  • Mucrorostrum yoroium Harris V.A. & Iwasaki, 1997

    MURRAMIA Huys, 2016 (syn. Murramia Harris V.A., 1994)

  • Murramia bicincta Harris V.A., 1994
  • Murramia magna (Harris V.A., 1994)
  • Murramia poorei (Walker-Smith, 2001)


    Subfamily Porcellidiinae Boeck, 1865

    PORCELLIDIUM Claus, 1860 (syn. Thyone Philippi, 1840 )

  • Porcellidium acutum Kim I.H. & H.S. Kim, 1997 (syn.Kensakia acuta (Kim I.H. & Kim H.S., 1997)
  • Porcellidium acuticaudatum Thompson I.C. & Scott A., 1903 (syn. Kensakia acuticaudata (Thompson I.C. & Scott A., 1903)
  • Porcellidium affinis Quidor, 1906 (syn. Porcellidium affine Quidor, 1906
  • Porcellidium aiiroa (Harris V.A. & Iwasaki, 1997) (syn. Kensakia acuta (Kim I.H. & Kim H.S., 1997)
  • Porcellidium akashimum Harris V.A. & Iwasaki, 1996
  • Porcellidium algoense Hicks, 1982
  • Porcellidium aoifuchidorum Harris V.A. & Iwasaki, 1997 (syn. Porcellidium wandoensis Kim I.H. & H.S. Kim, 1997
  • Porcellidium bicincta (Harris V.A., 1994) (syn. Murramia bicincta Harris V.A., 1994
  • Porcellidium bipartitum Kim I.H. & H.S. Kim, 1997 (syn. Dilatatiocauda bipartita (Kim I.H. & H.S. Kim, 1997)
  • Porcellidium brevicaudatum Thompson I.C. & Scott A., 1903 (syn. Acutiramus brevicaudatus (Thompson I.C. & Scott A., 1903)
  • Porcellidium brevicavum Kim I.H. & H.S. Kim, 1997
  • Porcellidium clavigerum Pesta, 1935 (syn. Clavigofera clavigera (Pesta, 1935)
  • Porcellidium dentatum Claus, 1860 (syn. Porcelloides tenuicaudus (Claus, 1860)
  • Porcellidium dilatatum Hicks, 1971 (syn. Dilatatiocauda dilatata (Hicks, 1971)
  • Porcellidium echinophilum Humes & Gelerman, 1962 (syn. Clavigofera echinophila (Humes & Gelerman, 1962)
  • Porcellidium erythrogastrum Harris V.A. & H.M. Robertson, 1994
  • Porcellidium erythrum Hicks, 1971
  • Porcellidium fasciatum Boeck, 1865
  • Porcellidium fimbriatum Claus, 1863 (syn.Porcellidium fimbriatum fimbriatum Claus, 1863)
  • Porcellidium fulvum Thomson G.M., 1883 (syn.Porcellidium clavigerum Pesta, 1935)
  • Porcellidium gamoi (Harris V.A. & Iwasaki, 1996) (syn. Kushia gamoi Harris V.A. & Iwasaki, 1996
  • Porcellidium hartmannorum Tiemann, 1978
  • Porcellidium hormosirii Harris V.A. & H.M. Robertson, 1994
  • Porcellidium kiiroum Harris V.A. & Iwasaki, 1996
  • Porcellidium laurencium Hicks, 1982 (syn. Clavigofera laurencia (Hicks, 1982)
  • Porcellidium lecanoides Claus, 1889 (syn. Porcellidium viride (Philippi, 1840)
  • Porcellidium londonarum Harris V.A., 1994
  • Porcellidium londonii Harris, 1994 (?)
  • Porcellidium magna Harris V.A., 1994 (syn. Murramia magna (Harris V.A., 1994)
  • Porcellidium naviculum Harris V.A. & H.M. Robertson, 1994
  • Porcellidium ocellum Harris V.A. & Robertson H.M., 1994
  • Porcellidium ofunatense Harris V.A. & Iwasaki, 1996
  • Porcellidium ovatum Haller, 1879 (syn. Porcellidium tenuicauda Claus, 1860)
  • Porcellidium paguri Ho, 1986 (syn. Kioloaria paguri (Ho, 1986)
  • Porcellidium peniculiferum Tiemann, 1978 (syn. Porcellidium tristanense Wiborg, 1964)
  • Porcellidium phyllosporum Harris V.A. & H.M. Robertson, 1994
  • Porcellidium planum Tiemann, 1977 (syn. Dilatatiocauda plana (Tiemann, 1977)
  • Porcellidium poorei Walker-Smith, 2001 (syn. Murramia poorei (Walker-Smith, 2001)
  • Porcellidium pulchrum Harris V.A. & H.M. Robertson, 1994
  • Porcellidium quinquelineatus (Harris V.A. & Robertson H.M., 1994) (syn. Acutiramus quinquelineatus Harris V.A. & Robertson H.M., 1994)
  • Porcellidium rastellum Harris V.A., 2014
  • Porcellidium roscoffensis (Bocquet, 1948)
  • Porcellidium ravanae Thompson I.C. & Scott A., 1903 (syn.Ravania ravanae (Thompson I.C. & Scott A., 1903)
  • Porcellidium rufolineatus (Harris V.A. & Robertson H.M., 1994) (syn. Acutiramus rufolineatus Harris V.A. & Robertson H.M., 1994)
  • Porcellidium rubrum Pallares, 1966
  • Porcellidium sarsi Bocquet, 1948 (syn. Porcellidium viride (Philippi, 1840)
    Porcellidium scutatum Claus, 1889 (syn. Porcelloides scutatus (Claus, 1889)
  • Porcellidium sesquimaculata Harris V.A., 1994 (syn. Acutiramus sesquimaculatus (Harris V.A., 1994)
  • Porcellidium similis Kim S.H. & Kim W., 1996 (syn. Acutiramus similis (Kim S.H. & Kim W., 1996)
  • Porcellidium tapui Hicks & Webber, 1983 (syn. Acutiramus tapui (Hicks & Webber, 1983)
  • Porcellidium tenuicauda Claus, 1860 (syn. Porcelloides tenuicaudus (Claus, 1860)
  • Porcellidium trisetosum Geddes, 1968 (syn. Geddesia trisetosa (Geddes, 1968)
  • Porcellidium tristanense Wiborg, 1964 (syn. Dilatatiocauda tristanensis (Wiborg, 1964)
  • Porcellidium ulvum Hicks, 1982 (syn. Clavigofera ulva (Hicks, 1982)
  • Porcellidium unicus Ummerkutty, 1970 (syn. Synurus unicus (Ummerkutty, 1970)
  • Porcellidium viridis (Philippi, 1840) (syn. Porcellidium viride (Philippi, 1840)
  • Porcellidium yoroium (Harris V.A. & Iwasaki, 1997) (syn. Mucrorostrum yoroium Harris V.A. & Iwasaki, 1997)
  • Porcellidium wandoensis Kim I.H. & H.S. Kim, 1997

    PORCELLOIDES Harris V.A., 2014

  • Porcelloides scutatus (Claus, 1889)
  • Porcelloides tenuicaudus (Claus, 1860)

    RAVANIA Harrris V.A., 2014

  • Ravania doliocauda Harris V.A., 2014
  • Ravania ravanae (Thompson I.C. & Scott A., 1903)
  • Ravania wellsi Harris V.A., 2014

    SYNURUS Harris V.A., 2014

  • Synurus ctenocheirus Harris V.A., 2014
  • Synurus unicus (Ummerkutty, 1970)

    TECTACIGULUM Huys, 2009

  • Tectacingulum nigrum (Harris V.A., 1994)
  • Tectacingulum tumidum (Harris V.A., 1994)


    Acutiramus Harris V.A. & H.M. Robertson, 1994 (= Kioloaria Harris V.A., 1994 (Junior homonym); Murramia Harris V.A., 1994 (= Murramia Huys, 2016; Thyone Philippi, 1840 (=Porcellidium Claus, 1860 (synonym)




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