CANCRINCOLIDAE Fiers, 1990 (syn. Ameiridae Boeck, 1865)


Th family include the genera Cancrincola Wilson, 1913 and Antillesia Humes,1958, formerly attributed to the family Ameridae, and the genus Abscondicola Fiers, 1990; all the species live in the branchial cavities of terrestrial decapods.


CANCRINCOLA Wilson, 1913

  • Cancrincola abbreviata Humes, 1957 [west Africa]
  • Cancrincola jamaicensis Wilson, 1913 [Jamaica]
  • Cancrincola longiseta Humes, 1957 [west Africa]
  • Cancrincola plumipes Humes, 1941 [USA]


    ANTILLESIA Humes,1958

  • Antillesia cardisomae Humes,1958 [central America]


    ABSCONDICOLA Fiers, 1990

  • Abscondicola humesi Fiers, 1990 [Nouvelle Guinee]


    NEOCANCRINCOLA Manè-Garzon & Sobota, 1974

  • Neocancrincola platensis Mane Garzon & Sobota, 1974




    SUPERFAMILY AEGISTHOIDEA Giesbrecht, 1892

    AEGISTHIDAE Giesbrecht 1892

    (syn.Cerviniidae Sars M., 1903)


    "The Aegisthinae is traditionally regarded as one of the three exclusively holoplanktonic families within the Harpacticoida. In contrast to the other families, Miraciidae and Clytemnestridae (now Peltidiidae), which primarily occur in the epipelagic zones of the oceans, aegisthids are typically mesopelagic or even bathypelagic. The recent discovery of a new primitive genus Andromastax Conroy-Dalton & Huys from hydrothermal vent sites at the Galapagos Rift raised the suspicion that the subfamily might also be distributed in the hyperbenthic environment (Conroy-Dalton & Huys, 1999). The latter, together with anchihaline caves and hydrothermal vents (and cold seeps) was recognized by Huys & Boxshall (1991) as one of the key habitats on the Earth’s surface in our continuous search for copepods of high phylogenetic significance.

    In response to the limited accessibility of these habitats, a number of sophisticated sampling techniques and sampling gear has been developed in recent years. For example, the Japan Marine Science and Technology Center (JAMSTEC) invested much effort in the design of suitable methods for sampling the benthopelagic organisms which inhabit the water overlying hydrothermal vents and cold seeps. Initially, benthopelagic sampling was performed by newly designed nets attached to the deep-sea submersible Shinkai 2000 (Kikuchi et al., 1990) but this method showed serious limitations in the quantitative estimation of benthopelagic plankton. This led to the alternative solution of attaching sampling gear to the highly successful Deep Tow System of JAMSTEC.

    The design of a multiple opening and closing plankton sampler (DT-MPS) by modification of Terazaki’s (1991) vertical multiple plankton sampler (ORI-VMPS) allowed benthopelagic sampling at a distance of 0.5 to 3m above the bottom. Using this sampling gear hyperbenthic copepods were collected in the cold seep area southeast off Hatsushima Island (Toda et al., 1995) and around the hydrothermal vent fields in the Okinawa ." [from: W. LEE & R. HUYS (2000)


    AEGISTHINAE Giesbrecht, 1892


    AEGISTHUS Giesbrecht 1891 (syn. Hensenella Dahl, 1895 )

  • Aegisthus aculeatus Giesbrecht 1891 [Atlantic Ocean]
  • Aegisthus atlanticus Wolfenden, 1902 (syn. Aegisthus mucronatus Giesbrecht, 1891
  • Aegisthus dubius Sars G.O., 1916 (syn. Aegisthus mucronatus Giesbrecht, 1891
  • Aegisthus longirostris Scott T., 1894 (syn. Aegisthus mucronatus Giesbrecht, 1891
  • Aegisthus mirabilis (Dahl F., 1895) (syn. Aegisthus mucronatus Giesbrecht, 1891
  • Aegisthus mucronatus Giesbrecht 1891 [Ireland, Atlantic Ocean; USA; marine, planktonic]

    AEGISTHIDAE incertae sedis

  • Aegisthus spinulosus Farran, 1905 [Ireland, Atlantic Ocean; marine, planktonic ]




    AMELOTES Por, 1969

  • Ameliotes mediterraneus (Soyer, 1970) (= Stratiopontotes mediterraneus Soyer, 1970

    ARCTICOCARELLA Kornev & Chertoprud, 2008

  • Arcticocarella reducta Kornev & Chertoprud, 2008

    ANDROMASTAX Conroy-Dalton & Huys, 1999

  • Andromastax muricatus Conroy-Dalton & Huys, 1998
  • Andromastax cephaloceratus Lee & Huys, 2000 [Japan; hydrothermal vents]

    NUDIVORAX Lee & Huys, 2000

  • Nudivorax todai Lee & Huys, 2000 [Japan; hydrothermal vents]

    SCABRANTENNA Lee & Huys, 2000

  • Scabrantenna islandica Apostolov, 2011 [Iceland]
  • Scabrantenna yooi Lee & Huys, 2000 [Japan; hydrothermal vents]

    JAMSTECIA Lee & Huys, 2000

  • Jamstecia terazakii Lee & Huys, 2000 [Japan; hydrothermal vents]


    CERVININAE Sars M., 1903


    CERVINIA Norman 1878 (syn. Neocervinia Huys, Møbjerg & Kristensen, 1997; Pseudocervinia Brodsky, 1963)

  • Cervinia bradyi Norman in Brady, 1878
  • Cervinia brevipes Brotskaya, 1963
  • Cervinia islandica Apostolov, 2011 [Iceland]
  • Cervinia itoi (Lee W & Yoo, 1998)
  • Cervinia langi Montagna, 1979
  • Cervinia magna Smirnov, 1946
  • Cervinia mediocauda Burgess, 1998
  • Cervinia philippinensis Huys, Møbjerg & Kristensen, 1997
  • Cervinia pilosa Lang, 1948
  • Cervinia plumosa
  • Cervinia synarthra Sars G.O., 1910 (syn, Cervinia synarthra synartha Sars G.O., 1910
  • Cervinia tenuicauda Brotskaya, 1963
  • Cervinia tenuiseta Brotskaya, 1963 (syn. Expansicervinia tenuiseta (Brotskaya, 1963)
  • Cervinia unisetosa Montagna, 1981

    SIPHONIS Mercado-Salas, Khodami & Martínez Arbizu, 2019

  • Siphonis aurreus Mercado-Salas, Khodami & Martínez Arbizu, 2019
  • Siphonis ruehlemanni Mercado-Salas, Khodami & Martínez Arbizu, 2019


    EXPANSICERVINIA Montagna, 1981 (syn. Brotskayaia Huys, Møbjerg & Kristensen, 1997)

  • Expansicervinia glacieria Montagna, 1981
  • Expansicervinia tenuiseta (Brotskaya, 1963)

    PARACERVINIELLA Brodsky, 1963

  • Paracerviniella denticulata Brotskaya, 1963

    Species of Cerviniinae (Cervinia Norman, 1878, Cerviniella Smirnov, 1946, Eucanuella T. Scott, 1901, Expansicervinia Montagna, 1981, Paracerviniella Brodskaya, 1963) o Brotskayaia Huys, Møbjerg & Kristensen, 1997 was synonymized with Expansicervinia Montagna, 1981 (Seifried, 2003). o Neocervinia Huys, Møbjerg & Kristensen, 1997 and Pseudocervinia Brodskaya, 1963, were synonymized with Cervinia Norman, 1878 (Seifried, 2003).


    CERVINIOPSINAE Brodskaya, 1963

    (Cerviniopsis Sars, 1909, Hemicervinia Lang, 1935, Herdmaniopsis Brodskaya, 1963, Pontostratiotes Brady, 1883, Stratiopontotes Soyer, 1970, Tonpostratiotes Itô, 1982.)


    CERVINIELLINAE Khodami, Mercado-Salas & Martínez Arbizu, 2020

    EUCANUELLA T. Scott, 1901

  • Eucanuella langi Por, 1964
  • Eucanuella longirostrata Itô Tat, 1983
  • Eucanuella reticulata Soyer, 1970
  • Eucanuella spinifera T. Scott, 1901

    FRISIA George & Viertel, 2020

  • Frisia magellanica George &Viertel, 2020 [Chile]

    HASE Corgosinho et al., 2018

  • Hase logomorphius Corgosinho et al., 2018
  • Hase talpamorphicus Corgosinho et al., 2018

    CERVINIELLA Smirnov, 1946 (syn. Askalonia Por, 1964 )

    "The genus Cerviniella Smirnov, 1946 was established by Smirnov (1946) for Cerviniella mirabilipes Smirnov, 1946, and originally placed in the family Cerviniidae Sars, 1903. However, subsequent phylogenetic studies have changed its taxonomic position. Seifried and Schminke (2003) reclassified Cerviniidae as a subfamily with the family Aegisthidae Giesbrecht, 1893, which includes two other subfamilies, Aegisthinae Giesbrecht, 1893, and Cerviniopseinae Brodskaya, 1963" (from: Cho et al., 2023)

  • Cerviniella abyssalis Apostolov, 2011 [Iceland]
  • Cerviniella arctica Kihara & Martinez-Arbizu, 2012 [Arctic]
  • Cerviniella bisegmenta Cho et al., 2023
  • Cerviniella bodini Coull, 1973
  • Cerviniella brodskayae Por, 1969
  • Cerviniella danae Kihara & Martinez-Arbizu, 2012 [Arctic]
  • Cerviniella gorbunovi Smirnov, 1946
  • Cerviniella hamata Coull, 1973
  • Cerviniella hitoshii Kihara & Martinez-Arbizu, 2012 [Arctic]
  • Cerviniella inermis Smirnov, 1946
  • Cerviniella langi Bodin, 1968
  • Cerviniella lagarderei Bodin, 1968
  • Cerviniella longifurcata Apostolov, 2011 [Iceland]
  • Cerviniella mirabilipes Smirnov, 1946
  • Cerviniella permixa Cho et al., 2023
  • Cerviniella peruana Becker, 1974
  • Cerviniella talpa (Por, 1964)

    Key to species of Cerviniella


    CERVINIOPSEINAE Brodskaya, 1963

    CERVINIOPSIS Sars, 1903

  • Cerviniopsis acutirostris Brotskaya, 1963
  • Cerviniopsis atlantica Apostolov, 2011 [Iceland]
  • Cerviniopsis clavicornis Sars G.O., 1903
  • Cerviniopsis curviseta Brotskaya, 1963
  • Cerviniopsis gorbunovi Smirnov, 1946
  • Cerviniopsis inermis Smirnov, 1946
  • Cerviniopsis intermedia Lang, 1936
  • Cerviniopsis langi Soyer, 1970
  • Cerviniopsis longicaudata Sars G.O., 1903
  • Cerviniopsis minutiseta Itô Tat, 1983
  • Cerviniopsis muranoi Itô Tat, 1983
  • Cerviniopsis obtusirostris Brotskaya, 1963
  • Cerviniopsis smirnovi Por, 1969
  • Cerviniopsis stylicaudata Lang, 1936
  • Cerviniopsis reducta Park et al., 2012 [Japan; marine]
  • Cerviniopsis spinosa Apostolov, 2011 [Iceland]

    HEMICERVINIA Lang, 1935

  • Hemicervinia ryforsi Lang, 1935 (syn. Hemicervinia stylifera (Thompson I.C., 1893)
  • Hemicervinia stylifera (Thompson I.C., 1893)

    HERDMANIOPSIS Brotskaya, 1963

  • Herdmaniopsis abyssicola Brotskaya, 1963
  • Herdmaniopsis malagassicus (Por, 1969) syn. Ameliotes malagassicus Por, 1969
  • Herdmaniopsis mediterraneus (Soyer, 1970) accepted as Stratiopontotes mediterraneus Soyer, 1970

    PONTOSTRATIOTES Brady, 1883

  • Pontostratiotes abyssicola Brady, 1883
  • Pontostratiotes acanthoferens Yamanaka, 1973
  • Pontostratiotes alatus Por, 1969
  • Pontostratiotes barnetti Dinet, 1978
  • Pontostratiotes bispinosa Apostolov, 2011 [Iceland]
  • Pontostratiotes cecilae Dinet, 1978
  • Pontostratiotes denticulatus Dinet, 1978
  • Pontostratiotes fontani Dinet, 1981
  • Pontostratiotes glaber Por, 1969
  • Pontostratiotes gladius Dinet, 1978
  • Pontostratiotes horrida Brotskaya, 1959
  • Pontostratiotes inermis Por, 1969
  • Pontostratiotes intermedia Apostolov, 2011 [Iceland]
  • Pontostratiotes ivelinae Apostolov, 2011 [Iceland]
  • Pontostratiotes lubricus Dinet, 1978
  • Pontostratiotes microserrulatus Por, 1969
  • Pontostratiotes minor Dinet, 1981
  • Pontostratiotes minutus Apostolov, 2011
  • Pontostratiotes pacificus Itô Tat, 1982
  • Pontostratiotes polaris Apostolov, 2011 [Iceland]
  • Pontostratiotes parapubescens Apostolov, 2011 [Iceland]
  • Pontostratiotes peruanus Becker & Schriever, 1979
  • Pontostratiotes polaris Apostolov, 2011
  • Pontostratiotes pori Dinet, 1978
  • Pontostratiotes pubescens Por, 1969
  • Pontostratiotes robustus Itô, 1982Itô , 1982
  • Pontostratiotes scotti Brotskaya, 1959
  • Pontostratiotes sixtorum Por, 1969
  • Pontostratiotes sixtorum mindannaensis Itô, 1982
  • Pontostratiotes unisetosus Itô , 1982
  • Pontostratiotes uxoris Dinet, 1978
  • Pontostratiotes vasconiensis Dinet, 1978
  • Pontostratiotes vitielloi Dinet, 1978
  • Pontostratiotes vivierae Dinet, 1978

    STRATIOPONTOTES Soyer, 1970

  • Stratiopontotes mediterraneus Soyer, 1970

    TONPOSTRATIOTES Itô, 1982

  • Tonpostratiotes tenuipedalis Itô,1982




    BALAENOPHILLIDAE Sars G.O., 1910

    BALAENOPHILLUS Aurivillius P.O.C., 1879 (syn. Harpactichechus Ortiz, Lalana & Torres-Fundora, 1992 )

  • Balaenophilus manatorum (Ortiz, Lalana & Torres-Fundora, 1992)
  • Balaenophilus umigamecolus Ogawa, Matasuzaki & Misaki, 1997
  • Balaenophilus unisetus Aurivillius P.O.C., 1879




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