CANTHOCAMPTIDAE Sars, 1906, emend. Monard, 1927, Lang, 1948


BRYOCAMPTUS Chappuis, 1928


In the following list, according to Kiefer (1978) and Dussart & Defaye (1990), the genus Bryocamptus s. l. Chappuis, 1928 with the subgenera Bryocamptus s.str. Chappuis, 1928, Rheocamptus , Borutzky, 1948, Limocamptus Chappuis, 1928 and Arcticocamptus Chappuis, 1928, will be considered. Wells (2007) do not report the the subgenus Rheocamptus, including the related species in the subgenus Bryocamptus.


BRYOCAMPTUS Chappuis, 1928 (subgenus BRYOCAMPTUS Chappuis, 1928)


  • Bryocamptus (Bryocamptus) aberrans Apostolov & Pesce, 1991 [Bulgaria; cave waters] [PDF]
  • Bryocamptus (Bryocamptus) abyssicola Borutzky & Okuneva, 1972 [Baikal lake]
  • Bryocamptus (Bryocamptus) baikalensis Borutzky, 1930
  • Bryocamptus (Bryocamptus) bogoriensis Kiefer, 1933 [India; fresh waters?]
  • Bryocamptus (Bryocamptus) borutzkyi Petkovski, 1969 (Asia, Afghanistan; fresh waters?]
  • Bryocamptus (Bryocamptus) bulbochaetus Borutzky, 1931 [Baikal lake]
  • Bryocamptus (Bryocamptus) campaneri Reid, 1994
  • Bryocamptus (Bryocamptus) chappuisi Borutzky, 1948 [Baikal lake]
  • Bryocamptus (Bryocamptus) cokeri Borutzky, 1948 [Baikal lake]
  • Bryocamptus (Bryocamptus) elaphoides Borutzky & Okuneva, 1972 [Baikal lake]
  • Bryocamptus (Bryocamptus) hiatus hiatus (Willey, 1925) [north America; moss, marsh]
  • Bryocamptus (Bryocamptus) hiatus australis (Coker, 1934) [north America; fresh waters]
  • Bryocamptus (Bryocamptus) hutchinsoni Kiefer, 1929 [USA; fresh waters]
  • Bryocamptus (Bryocamptus) incertus Borutzky, 1931 [Baikal lake]
  • Bryocamptus (Bryocamptus) intercalaris Shen & Tai, 1973 [China; cave waters]
  • Bryocamptus (Bryocamptus) kozhowi Borutzky & Okuneva, 1971 [Baikal lake]
  • Bryocamptus (Bryocamptus) longicaudatus Borutzky & Okuneva, 1972 [Baikal lake]
  • Bryocamptus (Bryocamptus) longifurcatus Borutzky, 1931 [Baikal lake]
  • Bryocamptus (Bryocamptus) longisetosus Borutzky, 1931 [Baikal lake]
  • Bryocamptus (Bryocamptus) minutus minutus (Claus, 1863) [cosmopolitan; fresh and brackish waters]
  • Bryocamptus (Bryocamptus) minutus var. bispinosa Lang, 1957 [Asia; fresh waters]
  • Bryocamptus (Bryocamptus) minutus schizodon (Mrazek, 1893)
  • Bryocamptus (Bryocamptus) minutus simplicidentata (Willey, 1934)
  • Bryocamptus (Bryocamptus) minutus vejodovskyformis Kiefer, 1934
  • Bryocamptus (Bryocamptus) mrazeki (Minkiewicz, 1916) [cenral Europe; fresh and brackish waters]
  • Bryocamptus (Bryocamptus) newyorkensis (Chappuis, 1927) [USA; fresh waters] [PDF]
  • Bryocamptus (Bryocamptus) pilosus Flossner, 1989
  • Bryocamptus (Rheocamptus) pygmaeus pygmaeus (Sars, 1863) [cosmopolitan, except Australia?; fresh waters, moss]
  • Bryocamptus (Rheocamptus) pygmaeus bulbochaetus Sterba, 1962 [Europe, Asia; fresh waters]
  • Bryocamptus (Bryocamptus) sinuatus Borutzky & Okuneva, 1972 [Baikal lake]
  • Bryocamptus (Bryocamptus) subarcticus (Wi8lley, 1925)
  • Bryocamptus (Bryocamptus) tarnogradskyi Borutzky, 1934 [Asia, Europe; caves, fresh waters]
  • Bryocamptus (Bryocamptus) tenuis Borutzky & Okuneva, 1971 [Baikal lake]
  • Bryocamptus (Bryocamptus) tuberculatus Borutzky, 1948 [Baikal lake]
  • Bryocamptus (Bryocamptus) umiatensis Wilson, 1958 [Alaska; fresh waters]
  • Bryocamptus (Bryocamptus) vejdovskyi vejdovskyi (Mrazek, 1893) [Europe, Asia, Japan ; fresh and brackish waters] [PDF]
  • Bryocamptus (Bryocamptus) vejdovskyi minutiformis Kiefer, 1934 [north America; fresh waters}
  • Bryocamptus (Bryocamptus) yohteiensis Ishida, 1997
  • Bryocamptus (Bryocamptus) washingtonensis Wilson, 1958 [north America; fresh waters]


    BRYOCAMPTUS Chappuis, 1928 (subgenus RHEOCAMPTUS)


  • Bryocamptus (Rheocamptus) albidus Okuneva, 1983 [Baikal lake]
  • Bryocamptus (Rheocamptus) alosensis Apostolov, 1998 [France; hyporheic waters]
  • Bryocamptus (Rheocamptus) aquaeductus aquaeductus Borutzky, 1934 [Europe; aqueducts]
  • Bryocamptus (Rheocamptus) aquaeductus var. petkovski Apostolov, 1971 [Bulgaria; springs]
  • Bryocamptus (Rheocamptus) balcanicus balcanicus (Kiefer, 1933) [Europe; cave, phreatic waters]
  • Bryocamptus (Rheocamptus) balcanicus f. babunae Petkovski, 1956 [Yugoslavia; ground waters] **
  • Bryocamptus (Rheocamptus) birsteini Borutzky, 1940 [URSS, cave waters]
  • Bryocamptus (Rheocamptus) bispinosus Borutzky, 1940 [URSS; cave waters]
  • Bryocamptus (Rheocamptus) borus Karanovic & Bobic, 1998 [Serbia; cave waters]
  • Bryocamptus (Rheocamptus) brevipes Borutzky, 1948 [Baikal lake]
  • Bryocamptus (Rheocamptus) crassipes Borutzky & Okuneva, 1972 [Baikal lake]
  • Bryocamptus (Rheocamptus) cristatus Borutzky & Okuneva, 1972 [Baikal lake]
  • Bryocamptus (Rheocamptus) dentatus Chappuis, 1937 [Italy, Spain, France; cave waters, wells]
  • Bryocamptus (Rheocamptus) denticulatus Borutzky & Okuneva, 1972 [Baikal lake]
  • Bryocamptus (Rheocamptus) gauthieri (Roy, 1924) [north America; moss]
  • Bryocamptus (Rheocamptus) innominatus Borutzky, 1940 [URSS, Azerbaidjan; cave waters]
  • Bryocamptus (Rheocamptus) littoralis Borutzky & Okuneva, 1972 [Baikal lake]
  • Bryocamptus (Rheocamptus) madarensis Apostolov, 1969 [Bulgaria; fresh waters]
  • Bryocamptus (Rheocamptus) marinus Gurney, 1927 [Suez; brackish waters]
  • Bryocamptus (Rheocamptus) mirus Petkovski & Karanovic, 1997 [Balkan Peninsula; Ohrid lake]
  • Bryocamptus (Rheocamptus) pirgos Apostolov, 1969 [Bulgaria; fresh waters]
  • Bryocamptus (Rheocamptus) pyrenaicus (Chappuis, 1923) [France, Spain; cave waters]
  • Bryocamptus (Rheocamptus) reductus Borutzky, 1948 [URSS, Azerbaidjan; cave waters]
  • Bryocamptus (Rheocamptus) rylovi Borutzky, 1931 [Baikal lake]
  • Bryocamptus (Rheocamptus) saxicola Borutzky & Okuneva, 1972 [Baikal lake]
  • Bryocamptus (Rheocamptus) spinulosus spinulosus Borutzky, 1934 [Asia; fresh waters]
  • Bryocamptus (Rheocamptus) spinulosus f. occidentalis Sterba, 1961 [Europe; fresh waters]
  • Bryocamptus (Rheocamptus) spinulosus f. triarticulata Sterba, 1968 [Asia; fresh waters?]
  • Bryocamptus (Rheocamptus) tauricus (Borutzky, 1930) [URSS; cave waters]
  • Bryocamptus (Rheocamptus) tatrensis (Minkiewicz, 1916) [Asia; fresh waters]
  • Bryocamptus (Rheocamptus) typhlops (Mrazek, 1893) [Europe; fresh waters,moss] [PDF]
  • Bryocamptus (Rheocamptus) unisaetosus Kiefer, 1930 * [Europe; cave waters]
  • Bryocamptus (Rheocamptus) zschokkei zschokkei (Schmeil, 1893) [cosmpolitan, except Australia?; fresh waters] [PDF]
  • Bryocamptus (Rheocamptus) zschokkei alleganiensis Coker, 1934 [USA; fresh waters]
  • Bryocamptus (Rheocamptus) zschokkei var. apilosus Sterba, 1955 [Europe; moss]
  • Bryocamptus (Rheocamptus) zschokkei caucasicus Kiefer, 1978
    [Asia; fresh waters] *
  • Bryocamptus (Rheocamptus) zschokkei f. kalinae Petkovski, 1956 [Yugoslavie;cave waters]
  • Bryocamptus (Rheocamptus) zschokkei komi Borutzky, 1962 [URSS; rivers]
  • Bryocamptus (Rheocamptus) zschokkei sinkiangensis Shen & Sung, 1965 [China; fresh waters?]
  • Bryocamptus (Rheocamptus) zschokkei triarticulata Kiefer, 1929 [France, URSS; ground waters] **
  • Bryocamptus (Rheocamptus) zschokkei yunnannsis Shen & Tai, 1964 [China; fresh waters]
  • Bryocamptus (Rheocamptus) weberi (Kessler, 1914) [Europe; fresh waters, wet moss]

    * syn. of B. (R.) tatrensis (Minkiewicz, 1916); according to Petkovski (1972) could be syn. of B. (R.) typhlops Mrazek, 1893)

    ** questionable taxa


    NO SUBGENERIC STATUS DEFINED

  • Bryocamptus lacustris Wells 2007
  • Bryocamptus pacificus Ishida, 1992 [Japan; streams]
  • Bryocamptus pilosus Flossner, 1989 [north America; lakes] [PDF]
  • Bryocamptus stouti Herding, 1958 [New Zealand, fresh waters]
  • Bryocamptus yohteiensis Ishida, 1997 [Japan; stream puddles]




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