PARASTENHELIIDAE Lang, 1944; sensu Willen (2000)


PARASTENHELIA Thompson & A. Scott, 1903

  • Parastenhelia bulbosa (Wells, 1963) [England; estuaries] [PDF]
  • Parastenhelia costata Pallares, 1982 [Argentina; marine]
  • Parastenhelia forficula Lang, 1936
  • Parastenhelia forficula littoralis Lang, 1934
  • Parastenhelia gracilis Brady, 1910 [Falkland, Kerguelen; marine]
  • Parastenhelia hornelli Thompson & A. Scott, 1903 [Black Sea, Sri Lanka, Antarctic (Livingston Island); sand banks with algae]
  • Parastenhelia jenkinsi [Australia; marine] Walker-Smith [PDF]
  • Parastenhelia megarostrum Wells, Hicks & Coull, 1982 [New Zealand; estuaries]
  • Parastenhelia minuta Pallares, 1982 [Argentina; marine]
  • Parastenhelia oligochaeta Wells & Rao, 1987 [Andaman, Nicobar islands; marine]
  • Parastenhelia ornatissima Por, 1964
  • Parastenhelia pyriformis Song et al., 2003 [Korea; sandy bottoms] [PDF]
  • Parastenhelia spinosa (Fischer, 1860) [Europe; north Africa; marine]
  • Parastenhelia spinosa f. bulgarica Apostolov, 1968 [Bulgaria;brackish waters]

    WORLD KEY TO PARASTENHELIA SPECIES [PDF]


    FOWEYA Gee, 2006 [PDF]


  • Foweya anglica (Norman & T. Scott, 1905) [England; marine] [PDF]
  • Foweya tertia (Kunz, 1975) [south Africa; tidal pools] [PDF]


    KARLLANGIA Noodt, 1964

    The genus Karllangia Noodt, 1964, formerly included in the family Ameiridae, has been recently removed to Parastenheliidae Lang, 1944 (Willen, 2000). The genus presently shows a scattered distribution, suggesting a circumtropical-subtropical distribution but, as Mielke (1994) suggest, "any statement on its biogeography is surely premature".

  • Karllangia arenicola arenicola Noodt, 1964 [Africa; marine interstitial] *
  • Karllangia arenicola bengalensis Wells & Rao, 1987 [Andaman, Nicobar; coastal waters]
  • Karllangia obscura Mielke, 1994 [Costa Rica; mesopsammon]
  • Karllangia ornatissima (Monard, 1935) [France, marine sand] [PDF]
  • Karllangia pulchra Mielke, 1994 [Costa Rica; mesopsammon]
  • Karllangia reducta (Apostolov, 1975) [Black sea] **
  • Karllangia tertia Kunz, 1975 ***

    * syn. Karllangia psammophila Wells, 1967 (Wells & Rao, 1987)

    ** incertae sedis according to Gee, 2006 [PDF]

    *** questionable taxon


    | KEY TO SPECIES OF KARLLANGIA | [PDF]





    RHIZOTHRICIDAE Por, 1986


    RHIZOTHRIX Brady & Robertson, 1875


    "The genus Rhizothrix Brady & Robertson, 1875 has a complex history. Brady & Robertson (1875) created the genus to accommodate R. curvata Brady & Robertson, 1875, the type species, but they did not include any description of the species. Later, Brady (1880) offered a detailed description of R. curvata. Pennak (1942) created the genus Adelopoda Pennak, 1942 and assigned it to the family Canthocamptidae Sars, 1906. Bodin (1997) was of the opinion that Adelopoda should have been assigned to the Cletodidae. Bozˇic´ (1953) considered Adelopoda a subgenus of Rhizothrix. Wells (1963) considered Adelopoda a synonym of Tryphoema Monard, 1926 and suggested that Tryphoema had to be considered a subgenus of Rhizothrix. Lang (1965) considered Rhizothrix and Tryphoema distinct, and reallocated the genus Adelopoda into the genus Tryphoema.

    Por (1986) allocated the genera Rhizothrix and Tryphoema to the family Rhizothricidae Por, 1986. with other members of the genus. A key to

    Currently, 12 species and subspecies are included within Rhizothrix. The type species of the genus, Rhizothrix curvata, was reported from British coasts.

    Later, Scott (1903) reported R. minuta (T. Scott, 1903) and R. gracilis (T. Scott, 1903) from Scotland. Wilson (1932) described R. tenella (Wilson, 1932) from the Atlantic coast of North America, and Lang (1936) reported R. scotti (Lang, 1936) from the Arctic Ocean. Since Lang’s (1948) monograph, six species and subspecies have been added: R. reducta Noodt, 1952; R. reducta noodti Galhano, 1970; R. pubescens Por, 1959; R. quadriseta Wells, 1967; R. spinosa Coull, 1971; and R. wilsoni Bodin, 1979. Most recently, Arlt (1983) reported R. cf. minuta from the Baltic Sea and discussed the differences observed between his R. cf. minuta and R. minuta" (from: Nam & Le, 2005).


  • Rhizothrix curvata Brady & Robertson, 1875 [Europe; marine sand]
  • Rhizothrix gracilis (T. Scott, 1903) [north Europe; marine sand]
  • Rhizothrix minuta (T. Scott, 1903) [north Europe; marine sand]
  • Rhizothrix pubescens Por, 1959 [Black Sea]
  • Rhizothrix quadriseta Wells,1967 [Mozambique; marine]
  • Rhizothrix reducta reducta Noodt, 1952 [Sylt island; marine]
  • Rhizothrix reducta noodti Galhano, 1970 [Portugal; marine interstitial]
  • Rhizothrix sejongi Jung Nam & Lee, 2005 [Korea; sandy beach] [PDF]
  • Rhizothrix scotti (Lang, 1936) [Arctic Ocean]
  • Rhizothrix spinosa Coull, 1971 [USA; marine]
  • Rhizothrix tenella (Wilson, 1932) [USA; Atlantic coast]
  • Rhizothrix wilsoni Bodin, 1979 [France; marine]


    | KEY TO THE SPECIES OF GENUS RHIZOTHRIX | [PDF]


    TRYPHOEMA Monard, 1926 [syn. ADELOPODA Pennak, 1942]

  • Tryphoema bocqueti (Bozic, 1953) [France; marine sand]
  • Tryphoema lusitanica (Wells & Clark, 1965) [Portugal; marine interstitial]
  • Tryphoema porca Monard, 1926
  • Tryphoema ramabula (Pennak, 1942) [USA; marine sand]
  • Tryphoema riedli Coull, 1971 [USA; marine sand]
  • Tryphoema scilloniensis (Wells, 1968) [Scilly island; marine]




    Page Updated May 07. 2008


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